Gudeodiscus phlyarius (Mabille 1887)

Figure 4.Shells of Vietnamese Gudeodiscus species. A Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), MNHN 24581 (syntype of Plectopylis phlyaria) B Gudeodiscus (Gudeodiscus) phlyarius, SMF 150125a (lectotype of Plectopylis moellendorffi) C Gudeodiscus (Gudeodiscus) cf. phlyarius, Vn10-41, coll. PGB D Gudeodiscus (Gudeodiscus) phlyarius, Vn09-06, coll. HE E Gudeodiscus (Gudeodiscus) phlyarius, NHMUK 1922.8.29.56 (holotype of Plectopylis gouldingi) F Gudeodiscus (Gudeodiscus) phlyarius, NHMUK 1922.8.29.57 (holotype of Plectopylis anterides). Photos: T. Deli (A), E. Neubert (B), B. Páll-Gergely (C, D) and H. Taylor (F). Scale represents 10 mm.

Figure 31.Inner wall of the vagina of Sicradiscus and Gudeodiscus species. A Sicradiscus mansuyi (Gude, 1908), (for locality see Figure 27) B Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn11-157 D Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 17).

Figure 35.SEM images of radulae of Gudeodiscus species. A, D, G, J, M show the middle part of the radula B, E, H, K, N show the central tooth and the first 2–3 pairs of laterals C, F, I, L, O show the marginals. A–C Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898) (for locality see Figure 19) D–F Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) G–I Gudeodiscus (Gudeodiscus) multispira (Möllendorff, 1883), China, Guangxi, Qingshan, Qingshan Zhen, Lipu Xian, ca 250 m, 24°26.189'N, 110°20.008'E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 12.11.2004. J–L Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Lạng Sơn Province, ca. km. 50 of road 1B, 10 km to Bình Gia, 21°53.911'N, 106°25.664'E, leg. Hemmen, Ch. & J., 01.04.2011. M–O Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), (for locality see Figure 23). All photos by B. Páll-Gergely.

Figure 43.Distribution of Gudeodiscus Páll-Gergely, 2013 species. Legends: filled triangle, top down: typical Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), filled triangle, top up: “Gudeodiscus phlyarius werneri Páll-Gergely, 2013” (synonym of phlyarius); empty triangle, top up: Gudeodiscus (Gudeodiscus) phlyarius populations showing transitional characters towards werneri in terms of shell shape; empty triangle, top down: atypical Gudeodiscus (Gudeodiscus) phlyarius; empty circle: Gudeodiscus messageri raheemi ssp. n., filled circle: Gudeodiscus (Gudeodiscus?) hemmeni sp. n. (in all localities it co-occurs with Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n.); circle with filled triangle in the middle: co-occurrence of Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n. and atypical Gudeodiscus (Gudeodiscus) phlyarius. The shaded area indicates the area inhabited by Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909) and “anterides”, “fallax” and “gouldingi”-like populations of Gudeodiscus (Gudeodiscus) phlyarius. Filled square indicates the position of Phong-Tho, the type locality of Plectopylis verecunda Gude, 1909 (synonym of Gudeodiscus phlyarius). Numbers 1–3 refer to atypical populations assigned to Gudeodiscus (Gudeodiscus) phlyarius. For explanation, see text.

Figure 16.Plot of shell height against shell width (diameter) for 122 adults of Plectopylis cf. anterides/gouldingi (MNHN 2012-2133, MNHN 2012-2156, partly MNHN 2012-2218), Plectopylis cf. fallax (Vn11-187, MNHN 2012-2132, partly MNHN 2012-2218) and Plectopylis cf. fallax var. major (MNHN 2012-2155) from northern Vietnam. Samples MNHN 2012-2155 and MNHN 2012-2156 originally belonged to the same sample.

Figure 5.Shells of Vietnamese Gudeodiscus species. A Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical “fallax var. major”), MNHN-IM-2012-2155 B Gudeodiscus (Gudeodiscus) phlyarius, NHMUK 1922.8.29.55 (holotype of Plectopylis verecunda) C Gudeodiscus (Gudeodiscus) phlyarius, NHMUK 1922.8.29.58 (holotype of Plectopylis fallax) D Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., Vn10-76, coll. PGB E Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n., NHMUK 20110370.1 (holotype) F Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909), NHMUK 1922.8.29.53 (holotype) G Gudeodiscus (Gudeodiscus) messageri messageri NHMUK 1922.8.29.54 (syntype of Plectopylis messageri var. minor). Photos: B. Páll-Gergely (A, D), H. Taylor (B–C, E–G). Scale represents 10 mm.

Figure 9.Shells (A–B) and apertural views (C–R) of Vietnamese Gudeodiscus, Sicradiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), MNHN 24586 (holotype?) B Gudeodiscus (Gudeodiscus?) suprafilaris, 2011/81 C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn11-156 D Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical “anterides/gouldingi”), MNHN-IM-2012-2164 E Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909), MNHN-IM-2012-2215 F Gudeodiscus (Gudeodiscus?) hemmeni Páll-Gergely & Hunyadi, sp. n., Vn10-103A G Gudeodiscus (Gudeodiscus?) anceyi (Gude, 1901), GS22 H Sicradiscus mansuyi (Gude, 1908), 20081116C I Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), 2011/81 J Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), Vn11-151 K–L Gudeodiscus (Gudeodiscus) dautzenbergi (Gude, 1901), Vn10-44 M–N Halongella schlumbergeri (Morlet, 1886), MAA3 O Halongella fruhstorferi (Möllendorff, 1901), Vn11-171 P Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), Vn10-120 Q Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), 20090515C R Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), 2011/81. All photos by B. Páll-Gergely except for Figure 9A (T. Deli). Scale represents 10 mm and refers to A and B.

Figure 10.SEM images of Gudeodiscus shells. A protoconch of Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., Vn12-104, coll. HE B protoconch of Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), Vn11-163, coll. HE C–D sculpture of Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn10-56, coll. HE E–F sculpture of Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical fallax specimen), Vn11-187, coll HE. Images: B. Páll-Gergely.

Figure 11.Parietal (A, C, E, G, I, K, M, O–Q, S, V) and palatal (B, D, F, H, J, L, N, R, T–U, W–X) plicae and lamellae of Sicradiscus and Gudeodiscus species. A–B Sicradiscus mansuyi (Gude, 1908), 20081116C (two different specimens) C–F Gudeodiscus (Gudeodiscus) anceyi (Gude, 1901) C–D figures in Gude (1901a) E MNHN-IM-2012-2263, F GS22 G–J Gudeodiscus (Gudeodiscus?) hemmeni Páll-Gergely & Hunyadi, sp. n. G–H 2012/62, spec.1 I–J 2012/62, spec.2 K–P Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical gouldingi and anterides shells) K–L Plectopylis gouldingi (after Gude 1909) M–N Plectopylis anterides (after Gude 1909) O–P MNHN-IM-2012-2153 Q–X Gudeodiscus (Gudeodiscus) phlyarius (typical fallax and fallax var. major shells) Q–R Plectopylis fallax (after Gude 1909) S MNHN-IM-2012-2157 T–U MNHN-IM-2012-2132 (2 different specimens) V–W MNHN-IM-2012-2155/6 (“var. major”, two different specimens), X Vn11-187. Inner view: D, L, N, R; Outer view: B, F, H, J, T, U, W, X.

Figure 12.Parietal (A, C, E, G–K, M, N, P–S, U, W) and palatal (B, D, F, L, P, T, V, X) plicae and lamellae of Gudeodiscus species. A–M Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) A–B Plectopylis phlyaria (after Gude 1901c) C–D Plectopylis moellendorffi (after Gude 1901c) E–F Vn10-49, G Vn09-24 H Vn10-56, I Vn9-16, spec.1 J Vn9-16, spec.2 K–M Plectopylis verecunda, MNHN 2012-2177 (3 different specimens) N–Q Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909) N–O Plectopylis messageri (after Gude 1909) P MNHN-IM-2012-2162, Q MNHN-IM-2012-2165 R–V Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n, R 20071116C, spec.1. S–T 20071116C, spec.2. U–V 20080509C W–X Vn10-104B. Inner views: B, D, F, O; Outer views: L, T, V, X.

Figure 21.Reproductive anatomy of Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical fallax). Locality information: Lào Cai Province, ca. 3 km SW of Nhà Văn Hóa, 22°25.513'N, 104°12.194'E, leg. Hemmen, Ch. & J., 04.10.2011. A–B “Specimen1” C “Specimen2”. Scale represents 1 mm.

Figure 22.Reproductive anatomy of Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887). Locality information: Lạng Sơn Province, ca. 10.6 km from Bình Gia to Lạng Sơn on road 1B, 21°53.639'N, 106°25.895'E, leg. Hemmen, Ch. & J., 01.04.2011. Scale represents 5 mm.

Figure 28.Inner walls of the penis of Gudeodiscus Páll-Gergely, 2013 species. A Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical fallax specimen, for locality see Figure 21) B Gudeodiscus giardi giardi (Fischer, 1898) (for locality see Figure 19) C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (for locality see Figure 22) D Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 17) E Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., 20080510A F–G Gudeodiscus (Gudeodiscus) villedaryi specimens collected at the same locality in two different dates: F November (2011/102) and G May (20090520A). All photos by B. Páll-Gergely.

Endoplon phlyaria (Mabille, 1887)

Plectopylis anterides Gude, 1909

Plectopylis anterides Gude, 1909

Endoplon phlyaria (Mabille, 1887)

Endoplon moellendorffi (Gude, 1901)

Endoplon fallax (Gude, 1909)

Endoplon moellendorffi (Gude, 1901)

Endoplon fallax (Gude, 1909)