Sphagnum

Sporophytes are uncommon in Sphagnum obtusum. This is a quite phenotypically variable species that warrants further investigation, which may result in taxonomic splitting. The strongly obtuse stem leaf should separate it from any similar species with which it occurs. Sphagnum mendocinum looks similar phenotypically but there appears to be no range overlap with S. obtusum. The tiny branch leaf pores, which may seem like no more than pinpricks in the cell surface, easily separate S. obtusum microscopically from other species of sect. Cuspidata.

Plants rather large, 8–12 cm high, pale green, tinged with yellowish brown, in loose or compact tufts. Stem cortex in 2–4 layers, hyaline cells large, thin-walled, without fibrils and pores; central cylinder pale yellowish brown, not clearly differentiated from cortical cells. Stem leaves 1.0–1.5 mm long, distantly set, triangular-ligulate, rounded, dentate or sometimes lacerate at the apex; borders narrow above, clearly widened below (ca. 1/3 the leaf base width); hyaline cells broadly rhomboidal, occasionally divided and fibrillose in the upper apex. Branches in fascicles of 4–5, with 2–3 spreading. Branch leaves 2.0–3.0 mm long, ovate-ligulate to lanceolate, undulate when dry, slightly involute near apex; hyaline cells linear-rhomboidal, densely fibrillose, with numerous, small, pore-like membrane thinnings in 1–2 rows on both surfaces; green cells in cross section narrowly isosceles-triangular, exposed on the dorsal surface, enclosed by hyaline cells or sometimes slightly exposed on the ventral surface. Dioicous. Sporophytes not seen.

Plants moderate to robust, weak-stemmed, yellow, yellowish brown to golden brown; capitulum varying from rounded, not 5-radiate and twisted to flat 5-radiate and straight branched. Stem pale green to pale brown; superficial cortex of weakly to moderately differentiated. Stem leaves triangular-lingulate, 0.9-1.3 mm; usually appressed; apex obtuse and often erose; hyaline cells efibrillose and nonseptate. Branches tapering or in more robust forms, frequently blunt, straight to arcuate, leaves slightly to moderately elongated at distal end. Branch fascicles with 2 spreading and 2 pendent branches. Branch stems green, with cortex enlarged with conspicuous retort cells. Branch leaves ovate to ovate-lanceolate; more than 1.8 mm; straight, stiff, not much undulate and reflexed to recurved; margins entire; hyaline cells on convex surface with a few end pores, but mostly numerous small to very small (often barely visible) pores or wall thinnings free from the commissures, on concave surface similar, but with pores generally fewer and larger; chlorophyllous cells triangular in transverse section, just reaching concave surface or slightly enclosed. Sexual condition dioicous. Spores 18-27 µm; both surfaces covered with rough, irregular verrucate plates of papillae, bifurcated Y-mark sculpture on distal surface; proximal laesura less than 0.5 spore radius.

Distribution: China, Japan, Russian Far East and Siberia, and Europe, and North America.

Habitat: in open bogs.

Sphagnum recurvum var. obtusum (Warnst.) Warnst., Hedwigia 23: 121. 1884.