Gudeodiscus giardi giardi

Figure 7.Shells of Vietnamese Gudeodiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus?) francoisi (Fischer, 1898), MNHN 24601 (holotype of Plectopylis bavayi) B Gudeodiscus (Gudeodiscus?) francoisi, MNHN 9945 (holotype of Plectopylis francoisi) C Gudeodiscus (Gudeodiscus?) francoisi, NHMUK 1922.8.29.51 (holotype of Plectopylis lepida) D Halongella fruhstorferi (Möllendorff, 1901), SMF 9258 (lectotype) E Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), MNHN IM-2010-12120 (syntype of Plectopylis congesta) F Gudeodiscus (Gudeodiscus) giardi giardi, NHMUK 1922.8.29.49 (syntype of Plectopylis congesta). Photos: T. Deli (A, B, E), H. Taylor (C, F) and E. Neubert (D). Scale represents 20 mm.

Figure 35.SEM images of radulae of Gudeodiscus species. A, D, G, J, M show the middle part of the radula B, E, H, K, N show the central tooth and the first 2–3 pairs of laterals C, F, I, L, O show the marginals. A–C Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898) (for locality see Figure 19) D–F Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) G–I Gudeodiscus (Gudeodiscus) multispira (Möllendorff, 1883), China, Guangxi, Qingshan, Qingshan Zhen, Lipu Xian, ca 250 m, 24°26.189'N, 110°20.008'E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 12.11.2004. J–L Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Lạng Sơn Province, ca. km. 50 of road 1B, 10 km to Bình Gia, 21°53.911'N, 106°25.664'E, leg. Hemmen, Ch. & J., 01.04.2011. M–O Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), (for locality see Figure 23). All photos by B. Páll-Gergely.

Figure 45.Living specimens of Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898) (A), Cao Bằng Province, Hòa An District, Nguyễn Huệ Commune, small hill just outside of Khau Trang Village, 22°33.510'N, 106°10.294'E, leg. Naggs, F. et al. 22.06.2011.; and Halongella schlumbergeri (Morlet, 1886) (B), Halong Bay area, Vietnam. Photos: F. Naggs.

Figure 42.Distribution of Gudeodiscus species. Legends: filled triangle: Gudeodiscus (Veludiscus) emigrans quadrilamellatus Páll-Gergely, 2013, empty circle: Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), filled circle: Gudeodiscus (Gudeodiscus?) francoisi (Fischer, 1899), semi filled circle: co-occurrence of the latter two species.

Figure 8.Shells of Vietnamese Gudeodiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), MNHN 9946 (syntype of Plectopylis giardi) B Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), SMF 9279 (lectotype of Plectopylis choanomphala) C Gudeodiscus (Gudeodiscus) villedaryi, NHMUK 1930.9.12.38 (holotype of Plectopylis villedaryi) D Gudeodiscus (Gudeodiscus) villedaryi, Vn11-152, coll PGB E Gudeodiscus (Gudeodiscus) dautzenbergi (Gude, 1901), MNHN 24603 (holotype) F Gudeodiscus (Gudeodiscus) dautzenbergi, MNHN 24602 (holotype of Plectopylis persimilis). Photos: T. Deli (A, E, F), S. Hof (B), H. Taylor (C) and B. Páll-Gergely (D). Scale represents 20 mm.

Figure 9.Shells (A–B) and apertural views (C–R) of Vietnamese Gudeodiscus, Sicradiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), MNHN 24586 (holotype?) B Gudeodiscus (Gudeodiscus?) suprafilaris, 2011/81 C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn11-156 D Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical “anterides/gouldingi”), MNHN-IM-2012-2164 E Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909), MNHN-IM-2012-2215 F Gudeodiscus (Gudeodiscus?) hemmeni Páll-Gergely & Hunyadi, sp. n., Vn10-103A G Gudeodiscus (Gudeodiscus?) anceyi (Gude, 1901), GS22 H Sicradiscus mansuyi (Gude, 1908), 20081116C I Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), 2011/81 J Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), Vn11-151 K–L Gudeodiscus (Gudeodiscus) dautzenbergi (Gude, 1901), Vn10-44 M–N Halongella schlumbergeri (Morlet, 1886), MAA3 O Halongella fruhstorferi (Möllendorff, 1901), Vn11-171 P Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), Vn10-120 Q Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), 20090515C R Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), 2011/81. All photos by B. Páll-Gergely except for Figure 9A (T. Deli). Scale represents 10 mm and refers to A and B.

Figure 13.Parietal (A, C, E, F, H, J, L, N, P, R, T, V, X) and palatal (B, D, G, I, K, M, O, Q, S, U, W, Y) plicae and lamellae of Gudeodiscus species. A–B Gudeodiscus (Veludiscus) emigrans emigrans (Möllendorff, 1901), holotype (after Páll-Gergely and Hunyadi 2013) C–D Gudeodiscus (Veludiscus) emigrans quadrilamellatus Páll-Gergely, 2013 (after Páll-Gergely and Hunyadi 2013) E–K Gudeodiscus (Gudeodiscus?) francoisi (Fischer, 1898) E–F holotype of Plectopylis lepida Gude, 1901 (after Gude 1901b), G MNHN-IM-2012-2311 H–I holotype of Plectopylis bavayi Gude, 1901 (after Gude 1901a) J–K Plectopylis francoisi (after Gude 1899b) L–U Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898) L Plectopylis giardi (after Gude 1899a) M Plectopylis giardi (after Gude 1899b) N–O Plectopylis congesta Gude, 1899 (after Gude 1899a); P–Q Vn10-69 R–S Vn10-59 T–U 2011/85 V–Y Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888) V–W holotype of Plectopylis (Endoplon) choanomphala Möllendorff, 1901 (after Gude 1901c) X–Y Vn10-47A. Inner views: D, F, I, K, M, O, Q, S, W, Y; Outer views: B, U.

Figure 19.Reproductive anatomy of Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898). Locality information: Cao Bằng Province, Quảng Uyên N, 206–207 cross, 430 m, 22°42.737'N, 106°27.223'E, leg. Hunyadi, A., 16.11.2011. Scales represents 5 mm (left) and 2 mm (right).

Figure 28.Inner walls of the penis of Gudeodiscus Páll-Gergely, 2013 species. A Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical fallax specimen, for locality see Figure 21) B Gudeodiscus giardi giardi (Fischer, 1898) (for locality see Figure 19) C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (for locality see Figure 22) D Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 17) E Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., 20080510A F–G Gudeodiscus (Gudeodiscus) villedaryi specimens collected at the same locality in two different dates: F November (2011/102) and G May (20090520A). All photos by B. Páll-Gergely.

Figure 29.Inner walls of male reproductive organs of Gudeodiscus and Halongella gen. n. species. A–B penis of Halongella schlumbergeri (Morlet, 1886), 20071122D C penis of Halongella fruhstorferi (Möllendorff, 1901) (for locality see Figure 25) D epiphallus of Gudeodiscus fischeri (Gude, 1901), (for locality see Figure 18) E epiphallus of Gudeodiscus giardi giardi (Fischer, 1898), (for locality see Figure 19) F penial caecum of Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) G penial caecum of Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n. (for locality see Figure 28E) H epiphallus of Halongella schlumbergeri (for locality see Figure 26) I Halongella fruhstorferi, (for locality see Figure 25) J spermatophore of Gudeodiscus (Gudeodiscus) fischeri, (for locality see Figure 17). All photos by B. Páll-Gergely.

Figure 30.Calcareous claws found in pockets on the inner penial wall of Gudeodiscus and Halongella gen. n. species. A–C, F Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888) (for locality see Figure 23) D Gudeodiscus giardi giardi (Fischer, 1898) (for locality see Figure 19) E Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 19) G–I Halongella schlumbergeri (Morlet, 1886) (for locality see Figure 29A–B). The claws in case of Gudeodiscus (Gudeodiscus) fischeri and Halongella schlumbergeri were too fragile for dissecting out, therefore drawings are presented. All images by B. Páll-Gergely.

Figure 32.Inner wall of the vagina of Halongella gen. n. and Gudeodiscus species. A–B Halongella fruhstorferi (Möllendorff, 1901), red circles indicate calcareous granules (for locality see Figure 25) C Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898) (for locality see Figure 19) D Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888) (for locality see Figure 24).