dcsimg
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Unresolved name

Animals

Metazoa

"""Ceratitis acicularis (Munro, 1969)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) acicularis (Munro)
Pterandrus acicularis Munro, 1969: 420.
Body length: 5.09 (4.80-5.30) mm; wing length: 5.68 (5.25-6.15) mm.

Male
Head:
Antenna yellow. First flagellomere 2-3 times as long as pedicel. Arista with moderately long rays; ventral rays somewhat shorter and sparser than dorsal rays, especially basally. Frons yellow; with short scattered setulae largely of same color as frons. Frontal setae well developed, occasionally less so. Face yellowish white. Genal seta and setulae dark, well developed. Thorax:
Postpronotal lobe white, without spot. Scutal pattern, ground color pale gray, with streaks and darker markings but without distinct spots or clearly defined stripes, except prescutellar white markings, separate but with pale area in between. Scapular setae dark. Scutellum grayish white, basally without spots, apically with three separate black spots, extending to basal half. Anepisternum completely pale, ventral margin narrowly dark brown; setulae pale except few dark setulae usually present along ventral margin. Legs:
Yellow except where otherwise noted; setation typical for subgenus, mainly dark. Foreleg:
femur anteriorly mainly brownish, only basal and ventral margin yellow, posteriorly largely brown, no distinct bush, only dispersed setulae; ventral setae black. Midleg:
femur anteriorly largely brown, distal end pale; ventrally with dark dispersed setulae, distally somewhat denser but without distinct feathering; tibia broadened; largely brown, with black feathering dorsally along distal 0.9 and ventrally along distal 0.8, anterior part covered by short dark setulae. Hindleg:
femur brown on basal 0.8; at apical 0.25 with longer setulae dorsally and ventrally. Wing:
Bands brownish. Interruption between marginal and discal bands near vein R1 clear and complete; discal band sometimes partly interrupted in discal cell; cubital band free; medial band absent; crossvein R-M opposite middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu straight. Abdomen:
Mostly Yellow. Tergites 2 and 4 largely pale gray. Tergite 3 with brownish black transverse band along posterior half. Tergite 5 along anterior margin with isolated brownish patches, posterior margin narrowly black.
Male
epandrium in lateral view with posterior lobe of lateral surstylus straight and elongate.
Female
As male except following characters:
Legs without feathering, yellow, except femora yellowish orange; forefemur posteroventrally with dark pilosity. Discal band not interrupted. Tergite 5 largely pale yellow across posterior half. Oviscape as long as, or longer than, preabdomen. Aculeus at least eight times as long as wide; tip with distinct apical indentation and lateral margin slightly sinuous.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis acicularis (Munro, 1969) in source PDF."""
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copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis aliena (Bezzi, 1920)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) aliena (Bezzi)
Pardalaspis aliena Bezzi, 1920: 231.
Body length: 4.02 (3.40-4.50) mm; wing length: 3.96 (3.65-4.35) mm.

Male
Head. Antenna yellow to yellow-orange. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons with short scattered hairs which are largely the same colour as frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum white, unspotted. Ground colour of mesonotum pale with orange tinge; spots pattern variable, sometimes spots confluent forming dark lateral stripes (in that case there are usually interrupted anteriorly of transverse suture). Chaetotaxy normal for subgenus. Scapular setae black, inner ones sometimes pale. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically, extending at most till halfway dorsal surface, sometimes limited to apical margin; basally without dark spots, only slight darker colouration. Subscutellum completely dark. Legs. Yellow; setation typical for subgenus, mainly pale, especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing. Bands with yellow-brown markings. Banding, marginal band with clear interruption after cell R1; cubital and medial bands joined with marginal band; crossvein r-m before middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Extension cup sometimes weakly sinoid. Abdomen. Pale yellow or more brownish. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Crossvein r-m rarely at middle. Oviscape shorter than abdominal terga 3-6 combined combined.
(Description after De Meyer, 1998)
See description of Ceratitis aliena (Bezzi, 1920) in source PDF."""
license
cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis andranotobaka Hancock, 1984"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) andranotobaka Hancock
Ceratitis andranotobaka Hancock, 1984: 285.
Body length: 4.40 mm; wing length: 4.50 mm.

Male
Head. Antenna yellow-orange. Third antennal segment two times as long as second segment. Arista with short hairs over entire length. Frons yellow to orange; with short scattered hairs which are distinctly darker than the frons; in lateral view not distinctly projecting forwards at antennal implant. Lower eye margin with darker colouration. Chaetotaxy normal for subgenus. Thorax. Postpronotum pale yellow, unspotted, only slightly paler than centre of mesonotum,. Ground colour of mesonotum pale with yellow-orange tinge. Chaetotaxy normal for subgenus. Scapular setae black. Two anepisternal bristles. Scutellum mainly yellow; with three black separate markings apically except at ventral margin where narrowly joining; basally two dark distinct spots. Subscutellum pale with three dark separate spots except along dorsal edge where touching. Legs. Yellow; setation typical for subgenus, mainly pale, especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing. Bands with markings extensively yellow. Banding, marginal band continuous; cubital band free; medial band joined with marginal band; crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Orange-brown. Setation and banding typical for subgenus.
Female
As male except for the following characters:
third antennal segment more orange coloured. Three frontal bristles. Inner scapular setae pale. Medial band interrupted in the middle. Oviscape as long as abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis andranotobaka Hancock, 1984 in source PDF."""
license
cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis anonae Graham, 1908"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) anonae Graham
Ceratitis anonae Graham, 1908: 114.
Body length: 5.06 (4.35-5.90) mm; wing length: 5.15 (4.45-5.75) mm.

Male
Head:
Antenna yellow. First flagellomere three times as long as pedicel. Arista with short to moderately long rays; ventral rays shorter and sparser than dorsal rays, especially basally. Frons pale, sometimes completely yellow, center yellow; with short scattered setulae distinctly darker than frons. Frontal setae well developed. Face white, sometimes yellowish white. Genal seta and setulae dark, well developed. Thorax:
Postpronotal lobe white, sometimes yellowish white; without spot. Scutal pattern:
ground color ash-gray; with streaks and darker markings but without distinct spots or clearly defined stripes except prescutellar white markings separate, sometimes with pale area in between. Scapular setae dark. Scutellum white, sometimes yellowish white, basally without spots, apically with three separate black spots, extending to basal 0.33, sometimes only to basal half. This has a strage meaning. The measurment should relate to apical 0.66 or 0.5 (global) I am not sure, the spots are apical spots reaching ‘forwards’ towards the basal margin. In some species they will reach the basal margin of the scutellum, in others only till halfway (‘basal half’ or ‘apical half’, whatever you choose), still in others till basal third. Referring this forward extension in relation to the apical length, sound a bit strange to me. I would therefore prefer to keep it in relation to the basal margin. However, if you feel very strongly about the apical aspect, you can still do a global change as such. Anepisternum on ventral 0.33-0.5 brown; setulae pale. Legs:
Yellow except where otherwise noted; setation typical for subgenus, mainly dark. Foreleg:
femur dark anteriorly along entire dorsal margin, posteriorly along dorsal 0.66, with poorly developed bush of dispersed long dark setulae along entire length, posterodorsal setulae longer; ventral setae dark. Midleg:
femur largely brownish black, anteriorly with silvery shine when viewed from certain angle, only distal end pale; ventrally with dark feathering along entire length, basally somewhat less dense; tibia broadened; largely brownish black with silvery shine when viewed from certain angle, with black feathering dorsally along distal 0.9 and ventrally along distal 0.8. Hindleg:
femur dark brown except distally, at apical 0.25 with longer setulae dorsally and ventrally. Wing:
Markings yellowish brown. Interruption between marginal and discal bands near vein R1 clear and complete; discal band often partly or completely interrupted in discal cell; cubital band free; medial band absent; crossvein R-M opposite middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu oblique anterobasally. Abdomen:
Mostly yellow. Border between tergite 1 and 2 narrowly black. Tergites 2 and 4 with pale gray band occupying almost entire tergite, at most narrowly yellow anteriorly. Tergite 3 with distinct brownish black band along posterior half; rarely interrupted medially. Tergite 5 with basal 0.33 brownish, usually divided medially into two spots; posteriorly narrowly brownish.
Male epandrium in lateral view with lateral surstylus curved, posterior lobe short.
Female
As male except following characters. Anepisternal pilosity on ventral 0.33 partly dark, especially centrally. Legs without feathering; femora yellow, forefemur posteriorly, and midfemur and hindfemur anteriorly on basal 0.66 often with brownish streaks; sometimes femora completely yellowish brown; forefemur posteroventrally with dark pilosity. Wing with discal band complete. Oviscape shorter than preabdomen. Aculeus at most six times as long as wide; tip with distinct apical indentation and lateral margin slightly sinuous.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis anonae Graham, 1908 in source PDF."""
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cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis antistictica Bezzi, 1913"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) antistictica Bezzi
Ceratitis stictica var. antistictica Bezzi, 1913: 20.
Body length: 4.35 mm; wing length: 4.15 mm.

Female
(Based on the original description, with additional notes based on the single BPBM specimen). Head. Antenna yellow-orange. Third antennal segment twice as long as second segment. Arista with short to medium long hairs over entire length (original description mentions quite long hairs). Frons with short scattered hairs which are the same colour as the frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Face yellow colour. Chaetotaxy normal for subgenus. Thorax. Postpronotum yellowish, more or less the same colour as centre of mesonotum (slightly less orange); with small darkish spot. Ground colour of mesonotum yellow-orange. Chaetotaxy normal for subgenus. Inner scapular setae pale, outer reddish. One anepisternal bristle. Scutellum mainly yellow, sometimes basally somewhat more whitish; with three black separate markings apically; basally two dark distinct spots (not clear in BPBM specimen since damaged by insect pin). Subscutellum pale with three dark separate spots except along dorsal edge where touching. Legs. Yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Ventral bristles on femora yellow. Wing. Bands with markings extensively yellow. Banding, marginal band continuous; cubital and medial bands free; discal band broad occupying most of basal part of wing; crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus. Oviscape shorter than abdominal terga 3-6 combined.
Male
No material seen. The original description mentions no apparent differences.
(Description after De Meyer, 1998)
See description of Ceratitis antistictica Bezzi, 1913 in source PDF."""
license
cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis argenteobrunnea Munro, 1935"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) argenteobrunnea Munro
Ceratitis (Pardalaspis) argenteobrunnea Munro, 1935: 312.
Body length: 5.08 (4.85-5.40) mm; wing length: 5.33 (5.25-5.4) mm.

Male
Head. Antenna yellow to yellow-orange. Third antennal segment three times as long as second segment. Arista with short hairs over entire length. Frons with short scattered hairs which are distinctly darker than the frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum yellowish, slightly less orange than centre of mesonotum; unspotted. Ground colour of mesonotum yellow-orange; mesonotal pattern, median and dorsocentral spots sometimes loosely connected with each other and with prescutellar spots (as in holotype). Chaetotaxy normal for subgenus. Scapular setae black. One anepisternal bristle. Scutellum mainly yellow, sometimes basally somewhat more whitish; with three black separate markings apically; basally two dark distinct spots. Subscutellum pale with three dark separate spots except along dorsal edge where touching. Legs. Yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing. Bands with markings extensively yellow. Banding, marginal band continuous; cubital band joined with discal band (not free as indicated in original description); medial band joined with marginal band; crossvein r-m before middle of discal cell. Crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen. Yellow or more brownish. Setation and banding typical for subgenus.
Female
As male except for the following characters:
lower eye margins with darker colouration. Thorax with inner scapular setae pale. Oviscape almost as long as abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis argenteobrunnea Munro, 1935 in source PDF."""
license
cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis brachychaeta Freidberg, 1991"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratitis) brachychaeta Freidberg
Ceratitis (Ceratitis) brachychaeta Freidberg, 1991: 169.
Body length: 3.97 (3.55-4.45) mm; wing length: 3.86 (3.40-4.45) mm.

Male
Head. Antenna yellow. Third antennal segment twice as long as second segment. Arista almost bare, at most with very short hairs on base. Frons pale yellow, in centre darker yellow; with short scattered hairs which are largely the same colour as frons. Frontal and orbital bristles yellow, ocellar black; lower orbital modified, flattened with bifurcated end, much shorter than arista; upper orbital slightly flattened and short but not modified otherwise (missing in most specimens studied, according original description shortly branched). Face yellow; lower eye margins sometimes darker yellow. Genal bristle pale, genal setulae pale, strongly developed. Postocellar and outer vertical bristles pale. Thorax. Postpronotum yellow-white, spot more brownish and weakly developed. Mesonotum ground colour black-brown, occasionally black, microtrichiae pattern silvery with ashgrey shine, spots black-brown or black except sutural yellow spots which are sometimes extended posteriorly as postsutural yellow lines, prescutellar yellow markings merged. Scapular setae pale. Scutellum yellow-white, basally with two dark merged spots, apically with three merged spots, incised to halfway. Anepisternum with lower half darker yellow but not distinct; also hairs on lower half somewhat darker yellow. Legs. Yellow; setation typical for subgenus, mainly pale especially on femora. Fore femur posteriorly with bush of longer orange-coloured hairs along entire length, posterodorsal hairs longer; ventral spines yellow-orange, anteroventral row of hairs short and yellow-orange. Hind femur with longer hairs dorsally and ventrally on apical fourth. Wing. Marginal band usually with clear and complete interruption, rarely with very weak and indistinct bridge; cubital band joined with discal band, sometimes bridge indistinct or cubital band free; medial band absent; crossvein r-m beyond middle of discal cell. Vein R  equal with crossvein r-m. Crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen. Yellow. Setation and banding typical of subgenus.
Female
As in male except for the following characters:
third antennal segment two to three times longer than second segment. Hairs on arista slightly more developed. Frons convex to flat, in lateral view very slightly projecting forwards at antennal implant. Orbitals not modified. All cephalic bristles dark, except postocellar and outer vertical. Legs without feathering. Wings, cubital band always free. Oviscape shorter than abdominal terga.
(Description after De Meyer, 2000)
See description of Ceratitis brachychaeta Freidberg, 1991 in source PDF."""
license
cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis bremii Guérin-Méneville, 1843"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pardalaspis) bremii Guérin-Méneville
Ceratitis bremii Guérin-Méneville, 1843: 199.
Body length: 5.88 (5.0-7.0) mm; wing length: 6.25 (5.5-7.0) mm.

Male
Head. Antennal segments yellow to yellow-orange. Third antennal segment twice as long as second segment. Arista with basal third yellow, otherwise dark. Frons pale to yellow ground colour, silvery over entire length; bare, no distinct hairs. Ocellar triangle dark. Face pale; with median yellow band. At antennal socket also with horizontal yellow band, sometimes both bands almost joining. Occiput moderately swollen below, pale whitish. Chaetotaxy normal for subgenus. Thorax. Ground colour of mesonotum pale with orange tinge; no distinct spots, three poorly defined darker stripes from anterior margin to dorsocentral setae, darker area between dorsocentral setae and prescutellar acrostichal setae, these markings not always distinct. Postpronotum same colour as centre of mesonotum. Chaetotaxy normal for subgenus. Two anepisternal bristles. Mesonotum with pale pilosity. Anepisternum completely pale pilose. Scutellum white basally, otherwise yellow with apical markings black; basally without dark spots, only slight darker coloration. Legs. Yellow; setation typical of subgenus, mainly pale especially on femora. Posterior row on fore femur pale. Wings. Bands with yellow markings extensive. Banding, setation and venation normal for subgenus. Marginal band continuous (in one specimen weakly interrupted); discal band joined with marginal band; cross-vein r-m before middle of discal cell; vein R  ending beyond cross-vein r-m. Abdomen. Pale yellow, with weakly defined spots. Pattern of spots and setation normal for subgenus.
Female
As male except for the following characters:
Frons yellower. Face with median band indistinct. Marginal band partly interruped at vein R  . Oviscape shorter than abdominal terga 3-6.
(Description after De Meyer, 1996)
See description of Ceratitis bremii Guérin-Méneville, 1843 in source PDF."""
license
cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis brucei Munro, 1935"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) brucei Munro
Ceratitis (Pardalaspis) brucei Munro, 1935: 308.
Body length: 5.40 mm; wing length: 5.75 mm.

Female
Head. Antenna yellow-orange. Antennal implant with a faint horizontal band (possibly discolouration). Third antennal segment twice as long as second segment. Arista missing in holotype. Frons with short scattered hairs which are largely the same colour or sligthly darker than frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum white, with black spot. Ground colour of mesonotum pale, with orange tinge. Chaetotaxy normal for subgenus. Scapular setae black. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically; basally with 2 dark, distinct spots. Subscutellum completely dark. Legs. Yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing. Bands with yellow-brown markings (yellow areas quite extensive but not predominant). Banding, marginal band continuous; cubital band free; medial band absent; crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen. Pale yellow-brown. Setation and banding typical for subgenus. Oviscape almost as long as abdominal terga 3-6 combined. Aculeus, tip in holotype exposed but broken; not dissected but apparently straight and simple, without serration or notches/appendices.
Male
Unknown.
(Description after De Meyer, 1998)
See description of Ceratitis brucei Munro, 1935 in source PDF."""
license
cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis caetrata Munro, 1949"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratitis) caetrata Munro
Ceratitis (Ceratitis) caetrata Munro, 1949: 499.
Body length: 4.02 (3.90-4.20) mm; wing length: 4.31 (4.00-4.65) mm.

Male
Head. Antenna yellow. Third antennal segment twice as long as second segment. Arista with short hairs. Frons yellow, inflated above antennal implant; almost bare, at most with few short scattered hairs which are largely the same colour as frons. Ocellar and orbital bristles yellow; lower orbital modified, apical end dark and fan shaped, at least twice as long as arista; upper orbital much reduced, weakly developed. Frontal bristles absent. Face yellow-white. Genal bristle pale, genal setulae dark reddish, weakly developed. Postocellar and outer vertical bristles pale. Thorax. Postpronotum pale yellow, with distinct black spot. Mesonotum:
ground colour orange; microtrichiae pattern silvery with orange shine, spots brown or black-brown; sutural spots whitish, small; prescutellar whitish markings separate. Scapular setae pale. Scutellum yellow-white, basally with two dark separate spots, apically with three merged spots, incisions variable, at least till halfway and sometimes almost reaching apical bristles. Anepisternum completely pale. Legs. Yellow; setation typical for subgenus, mainly pale especially on femora. Fore femur posteriorly with bush of longer orange-coloured hairs along entire length, posterodorsal hairs longer; ventral spines yellow-orange; anteroventral row of hairs short and yellow. Hind femur with longer hairs dorsally and ventrally on apical fourth. Wing. Marginal band with clear and complete interruption; cubital band free; medial band absent; crossvein r-m at middle of discal cell. Vein R  beyond crossvein r-m. Crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As in male except for the following characters:
Third antennal segment two to three times as long as second segment; yellow-orange. Frons yellow-orange, only slightly inflated; two frontal bristles present; short hairs more developed and distinctly darker than frons. Ocellars dark (frontal and orbitals presumably also but missing in specimens studied). Orbitals not modified (according original description). Genal setulae strongly developed. Mesonotal pattern sometimes much darker, more like in C. capitata. Crossvein dm-cu posteriorly more inwards. Legs without feathering. Oviscape shorter than abdominal terga.
(Description after De Meyer, 2000)
See description of Ceratitis caetrata Munro, 1949 in source PDF."""
license
cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis capitata (Wiedemann, 1824)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratitis) capitata (Wiedemann)
Tephritis capitata Wiedemann, 1824: 55.
Body length: 4.00 (3.45-4.60) mm; wing length: 4.12 (3.65-4.55) mm.

Male
Head. Antenna yellow or more orange; first and second segment and base of third segment sometimes darker. Third antennal segment twice as long as second segment. Arista with short hairs, mainly on base and only distinct dorsally. Frons convex to flat; yellow, sometimes with darker orange or orange-brown patches including darker band near antennal implant, occasionally with faint silvery shine; with short scattered hairs which are largely the same colour as frons. Frontal and ocellar bristles black; lower orbital modified, stem pale and shorter than arista with apical end dark and diamond-shaped; upper orbital weakly developed, black. Face yellow-white. Genal bristle pale, genal setulae pale or reddish, weakly developed. Postocellar and outer vertical pale. Thorax. Postpronotum white, with distinct black spot. Mesonotum:
ground colour black, microtrichiae pattern silvery with ashgrey shine, spots black except sutural white spots, prescutellar white markings merged. Scapular setae pale. Scutellum yellow-white, basally with two dark spots, separate or narrowly touching, apically with three merged spots, only slightly incised. Anepisternum pale with lower half darker yellow, pilosity variable but at least partly dark in lower half. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Fore femur posterodorsally with bush of longer orange-coloured hairs along entire length, basally these hairs darker red or brown but not distinct black; posteriorly hairs much shorter; ventral spines yellow-orange; anteroventral row of hairs short and yellow-orange. Hind femur with longer hairs dorsally and ventrally on apical fourth. Wing. marginal band usually with clear and complete interruption, occasionally narrowly or partly touching; cubital band free; medial band absent; crossvein r-m at or near middle of discal cell. Vein R  beyond or equal with crossvein r-m. Orientation crossvein dm-cu variable. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As in male except for the following characters:
Third antennal segment in general darker than in male. Frons sometimes with darker hairs; darker patches never as outspoken as in male. Orbitals not modified, well developed. Genal setulae darker and strongly developed. Anepisternum without darker pilosity. Legs without feathering; ventral spines on fore femur sometimes partially dark. Oviscape shorter than abdominal terga.
(Description after De Meyer, 2000)
See description of Ceratitis capitata (Wiedemann, 1824) in source PDF."""
license
cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis catoirii Guérin-Méneville, 1843"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratitis) catoirii Guérin-Méneville
Ceratitis catoirii Guérin-Méneville, 1843: 197.
Body length: 5.25 (4.65-5.55) mm; wing length: 5.26 (4.70-5.55) mm.

Male
Head. Antenna pale yellow. Third antennal segment twice as long as second segment. Arista almost bare, at most with very short hairs on base. Frons pale yellow; with short scattered hairs which are largely the same colour as frons. Frontal bristles yellow, weakly developed; ocellar and orbital bristles pale or dark; lower orbital modified, about as long as arista, apical end white and broadly spatulate shaped; upper orbital weakly developed. Face yellow-white. Genal bristle pale; genal setulae pale, weakly developed. Postocellar and outer vertical pale. Thorax. Postpronotum yellow-white, with distinct black spot. Mesonotum:
ground colour black, microtrichiae pattern silvery with ashgrey shine, spots black except sutural yellow-white spots, prescutellar yellow or yellow-white markings separate or merged. Scapular setae pale. Scutellum yellow-white, basally with two dark merged spots, apically with three merged spots, only slightly incised. Anepisternum pale yellow, lower half darker yellow but indistinct, hairs in lower half also slightly darker yellow. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Fore femur posteriorly with bush of longer orange-coloured hairs along entire length, posterodorsal hairs longer. Anteroventrally row of longer hairs, as long as posterodorsal hairs; ventral spines yellow-orange. Mid femur with row of long hairs anteroventrally. Hind femur with longer hairs dorsally and ventrally on apical fourth. Wing. Marginal band with clear and complete interruption; cubital band joined with discal band; medial band free; crossvein r-m at middle of discal cell. Vein R  beyond or equal with crossvein r-m. Crossvein dm-cu straight. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As in male except for the following characters:
Third antennal segment darker, yellow to yellow-orange. Hairs on arista slightly more developed. Frons yellow, occasionally more palish; with short hairs which are distinctly darker than frons. All cephalic bristles dark, except postocellar and outer vertical. Orbitals not modified, frontals well developed. Genal bristle pale or dark; genal setulae dark, strongly developed. Sometimes few dark hairs in lower half of anepisternum. Scapular setae rarely black. Legs in general with darker pilosity; no feathering; ventral spines on fore femur black. Wings, medial band occasionally touching marginal. Crossvein dm-cu posteriorly more inwards than anteriorly. Oviscape shorter than abdominal terga.
(Description after De Meyer, 2000)
See description of Ceratitis catoirii Guérin-Méneville, 1843 in source PDF."""
license
cc-by-nc-sa-3.0
copyright
"""Marc de Meyer, MRAC"""
editor
Marc de Meyer

"""Ceratitis colae Silvestri, 1914"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) colae Silvestri
Ceratitis colae Silvestri, 1913: 63.
Body length: 4.95 (4.30-5.50) mm; wing length: 5.19 (4.25-5.90) mm.

Male
Head:
Antenna yellowish orange to orange. First flagellomere with dorsal margin sometimes brown; 2-3 times as long as pedicel. Arista with moderately long rays; ventral rays shorter and sparser than dorsal rays, especially basally. Frons convex or flat; yellow to orange; with short scattered setulae distinctly darker than frons. Frontal setae well developed. Face yellow; gena brown, partly continued along ventral margin of face but not in median part; mouthparts with apex partly darkened but not completely black (as in lepida). Genal seta and setulae dark, well developed. Thorax:
Postpronotal lobe white, without spot. Mesonotum:
ground color dark gray with silvery shine; with streaks and darker markings but without distinct spots or clearly defined stripes, except prescutellar yellow markings, which are separate. Scapular setae dark. Scutellum white, basally without spots, apically with three separate black spots, extending to half-way. Anepisternum on ventral half brownish black; setulae on ventral half black, ventral to anepisternal seta with longer black setulae. Legs:
Yellow except where otherwise noted; setation typical for subgenus, mainly dark. Foreleg:
coxa brown, occasionally almost black; femur anteriorly with contrasting black/white pattern, basally pale yellow; white spot turns silverly when viewed from certain angle, anteroventrally with bush of short dark setulae in median half; posteriorly slightly darkened along dorsal and ventral margin; with dispersed and poorly developed bush of long dark setulae along entire length, posterodorsal setulae longer; ventral setae dark; tibia anteriorly slightly darkened. Midleg:
femur anteriorly yellow to yellowish brown; anteroventrally with conspicuous silvery patch, distally with inconspicuous one; ventrally with dark feathering along entire length except for interruption (6-7 setae wide) in median part; tibia broadened; yellow, silvery shine when viewed from certain angle; with black feathering dorsally along distal 0.8 and ventrally along distal 0.66, anteriorly with dispersed short setulae, in front of dorsal row forming an irregular second one. Hindleg:
femur at apical 0.25 with longer setulae dorsally and ventrally. Wing:
Bands brown or yellowish brown. Interruption between marginal and discal bands near vein R1 clear and complete; cubital band free; medial band absent; crossvein R-M proximal to middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu oblique anterobasally, or straight. Abdomen:
Mostly yellow. Tergite 1 with small brown patches at posterior margin. Tergites 2 and 4 with silvery transverse band along posterior half. Tergite 3 with brown transverse band along posterior 0.33, sometimes interrupted in middle. Tergite 5 with yellowish brown patches along anterior part and weak silvery band posteriorly. Sometimes abdomen with general darker brown appearance. Setation and banding typical for subgenus.
Male epandrium in lateral view with posterior lobe of lateral surstylus elongate, apical end curved.
Female
As male except for the following characters:
Legs without feathering; yellow, femora darker; forefemur posteroventrally with dark pilosity. Wing usually with darker banding. Abdominal tergite 5 with distinct silvery band posteriorly. Oviscape almost as long as preabdomen. Aculeus at least eight times longer than wide; tip with distinct apical indentation and lateral margin slightly sinuous.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis colae Silvestri, 1914 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis connexa (Bezzi, 1926)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) connexa (Bezzi)
Hoplolophomyia connexa Bezzi, 1926: 283.
Body length: 7.31 (6.85-7.85) mm; wing length: 7.18 (7.00-7.50) mm.

Male
Head. Antenna yellow to yellow-orange. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons yellow with lower half shining silvery; with short scattered hairs which are largely the same colour as frons; usually convex, sometimes flat, in lateral view slightly projecting forwards at antennal implant. Lower eye margin without distinct patch, at most slightly darker yellow colouration. Chaetotaxy normal for subgenus, except only one frontal bristle developed, sometimes two additional very small ones. Thorax. Postpronotum sligthly paler yellow than mesonotum, almost white; unspotted. Ground colour of mesonotum pale with yellow tinge; sometimes very pale and thereby obscuring white patches and lines. Chaetotaxy normal for subgenus. Scapular setae black. Two anepisternal bristles, rarely three. Scutellum white basally, otherwise yellow with three black separate markings apically; basally without dark spots. Subscutellum with three separate spots, same position as scutellar spots. Legs yellow; setation typical for subgenus, mainly pale especially on femora, partly dark reddish. Posterior and posterodorsal rows on fore femur pale, sometimes more reddish. Wing bands with extensive yellow markings. Banding, marginal band continuous; cubital band joined with discal band, medial with marginal band; crossvein r-m at middle of discal cell (occasionally just before middle). Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow ground colour. Anterior margin of fourth abdominal tergite with four and fifth tergite with two less distinct spots; no distinct banding.
Female
As male except for the following characters:
Antenna yellow. Three frontal bristles, of approximately equal length. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis connexa (Bezzi, 1926) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis contramedia (Munro, 1937)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) contramedia (Munro)
Pardalaspis contramedia Munro, 1937: 2.
Body length: 5.14 (4.75-5.50) mm; wing length: 5.29 (4.20-6.75) mm.

Male
Head. Antenna yellow-orange. Third antennal segment three times as long as second segment. Arista bare. Ocellar bristles well developed but short, at most 1.5 times as long as ocellar triangle. Frons with short scattered hairs which distinctly darker than frons; more flattened, not distinctly convex, in lateral view distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum more or less concolorous with centre mesonotum, only slightly less orange, unspotted. Ground colour of mesonotum pale with orange tinge. Chaetotaxy normal for subgenus. Scapular setae pale. One anepisternal bristle. Scutellum yellow with three apical black markings which are separated except narrowly joined along apical margin; basally without dark spots, only slight darker colouration. Subscutellum with two spots. Legs yellow; setation typical for subgenus, with mixed pale and darker hairs. Posterior and posterodorsal rows on fore femur with pale bristles except apical half of posterior row with dark bristles. Wing bands with yellow-brown markings; bands not very clear in some specimens especially not discal and cubital bands. Banding, marginal band continuous; cubital band free; medial band absent; crossvein r-m well before middle of discal cell, usually at basal third. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Pale yellow or more brownish, with darker coloured spots (spots not as outspoken as in Pardalaspis species). Otherwise setation and banding typical for subgenus.
Female
As male except for the following characters:
Oviscape at least as long as abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis contramedia (Munro, 1937) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis cornuta (Bezzi, 1924)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratitis) cornuta (Bezzi)
Pterandrus cornutus Bezzi, 1924: 478.
Body length: 4.33 (4.00-4.65) mm; wing length: 4.46 (4.20-4.70) mm.

Male
Head. Antenna yellow. Third antennal segment twice as long as second segment. Arista with short hairs, slightly longer than in other Ceratitis. Frons pale yellow; with short scattered hairs which are largely the same colour as frons. Frontal, ocellar and orbital bristles yellowish red, upper orbital sometimes dark; lower orbital with variable tip shape:
pointed, or flattened; frontal bristles weakly developed. Face pale yellow. Genal bristle pale; genal setulae pale, weakly developed. Postocellar and outer vertical bristles pale. Thorax. Postpronotum white, with distinct black spot. Mesonotum:
ground colour orange, occasionally more orange-brown, microtrichiae pattern silvery with orange shine, mesonotal spots more brownish; sutural yellow spots occasionally extended as postsutural lines, two separate prescutellar yellow lunules. Scapular setae pale. Scutellum yellow-white, basally with two dark separate spots, apically with three merged spots, incisions deep, almost reaching apical bristles. Anepisternum with lower half darker yellow but not distinct, pilosity in lower half darker yellow. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Fore femur posteriorly with weakly developed bush of longer orange-coloured hairs along entire length, posterodorsal hairs longer; ventral spines yellow-orange; anteroventrally without row of hairs. Mid femur with few longer hairs basally along ventral side but no distinct row. Hind femur with longer hairs dorsally and ventrally on apical fourth. Wing, brownish marking slightly more reduced than in other Ceratitis species. Banding, marginal continuous or with partial interruption (rarely completely interrupted); cubital and medial bands free; crossvein r-m at middle of discal cell. Vein R  beyond or equal with crossvein r-m. Crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As in male except for the following characters:
Frons convex to flat, slightly darker than in male. Frontal, ocellar and orbital bristles all dark; orbitals not modified, frontal bristles well developed. Genal bristles pale or dark. Postocellar and outer vertical bristles dark. Genal bristle dark or pale; genal setulae dark, strongly developed. Outer scapular black. Incisions apical scutellar spots variable but not as deep as in male; basal spots narrowly touching. Legs without feathering. Oviscape shorter than abdominal terga.
(Description after De Meyer, 2000)
See description of Ceratitis cornuta (Bezzi, 1924) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis cosyra (Walker, 1849)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) cosyra (Walker)
Trypeta cosyra Walker, 1849: 1042.
Body length: 4.43 (3.35 - 5.40) mm; wing length: 4.17 (3.40-5.20) mm.

Male
Head. Antenna yellow-orange. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons with short scattered hairs which are distinctly darker than or same colour as frons; more flattened, not distinctly convex, in lateral view slightly projecting forwards at antennal implant. Lower eye margin with slightly darker marking. Chaetotaxy normal for subgenus, bristles dark reddish to black. Thorax. Postpronotum white, with black spot. Ground colour of mesonotum pale with orange tinge; mesonotal pattern variable especially spots at mesal end of suture and prescutellar spots variable in size and colouration, anterior supra-alar spots usually continuous. Chaetotaxy normal for subgenus. Scapular setae pale. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically; basally usually with two separate dark spots, sometimes spots not outspoken, and only present as slightly brown patches. Subscutellum pale with three dark separate spots except along dorsal margin where touching. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Ventral spines on fore femur yellowish or black. Wing bands with markings extensively yellow; banding sometimes faint. Banding, marginal band continuous; cubital band free; medial band absent; crossvein r-m before middle of discal cell. Crossvein dm-cu position variable. Abdomen. Pale yellow or more brownish. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis cosyra (Walker, 1849) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis cuthbertsoni (Munro, 1936)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pardalaspis) cuthbertsoni (Munro)
Pardalaspis cuthbertsoni Munro, 1936: 42.
Pardalaspis cuthbertsoni var. nigrotertius Munro, 1939: 4.
Body length: 7.94 (7.5-8.5) mm; wing length: 8.0 (7.0-9.0) mm.

Male
Head. Antennal segments orange to dark orange. Third antennal segment at most twice as long as second segment. Arista basal part orange, apically darker. Frons orange to dark red in ground color, lower third silvery as a distinct band; dispersed short hairs, only slightly darker than ground colour. Ocellar triangle dark. Face orange-red; with median whitish band. Occiput moderately swollen below, pale whitish. Chaetotaxy normal for subgenus. Thorax. Ground colour of mesonotum brownish (dark grey in var. nigrotertius); no distinct spots, three poorly defined darker stripes from anterior margin to dorsocentral setae, darker area between dorsocentral setae and prescutellar acrostichal setae especially around the latter, these markings not always distinct and often interrupted. Postpronotum same colour as centre of mesonotum (in var. nigrotertius paler). Chaetotaxy normal for subgenus. Mesonotum with pale pilosity. Anepisternum with pale pilosity except upper margin with darker hairs. Two anepisternal bristles. Scutellum yellow with apical markings black; basally without dark spots, only faintly darker yellow coloration. Legs dark yellow to orange brown; setation typical of subgenus. Front femur with mainly pale pilosity, posterior row dark Wings with brownish bands, yellow markings strongly reduced. Banding, setation and venation normal for subgenus. Marginal band interrupted at vein R  ; discal band joined with marginal band but interrupted in discal cell (in some specimens interruption is very weak and bands appear continuous); cross-vein r-m usually before middle of discal cell, sometimes at middle; vein R  usually ending opposite or beyond cross-vein r-m. Abdomen. grey-brown, with clearly defined spots. Pattern of spots and setation normal for subgenus.
Female
As male except for the following characters:
Frons yellow to yellow-orange, lower margin distinctly paler. Face wholly yellow. Markings on mesonotum less clearly defined. In var. nigrotertius position of R  variable, either before or beyond cross-vein. Oviscape very long, about twice as long as abdominal terga 3-6; dark orange to brown.
(Description after De Meyer, 1996)
See description of Ceratitis cuthbertsoni (Munro, 1936) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis discussa Munro, 1935"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) discussa Munro
Ceratitis (Pardalaspis) discussa Munro, 1935: 307.
Body length: 5.42 (5.00-5.85) mm; wing length: 4.93 (4.40-5.40) mm.

Male
Head. Antenna yellow-orange. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons with short scattered hairs which are same colour as frons; more flattened, not distinctly convex, in lateral view slightly projecting at antennal implant. Lower eye margin at most with slightly darker yellow marking. Chaetotaxy normal for subgenus, dark reddish. Thorax. Postpronotum white, with black spot. Ground colour of mesonotum pale with yellow tinge; mesonotum:
spots at mesal end of transverse suture sometimes smaller and/or only darkish brown, anterior supra-alar spots usually at least partly interrupted. Chaetotaxy normal for subgenus. Scapular setae pale. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically; basally without dark spots, at most darker yellow colouration. Subscutellum pale with three dark separate spots, sometimes touching along dorsal margin. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Ventral spines on fore femur pale. Wing bands with markings extensively yellow. Banding, marginal band continuous; cubital band free; medial band absent; crossvein r-m before middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Pale yellow. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Third antennal segment somewhat longer, almost three times as long as second segment. Frons with scattered hairs which are darker than frons. Crossvein r-m sometimes at middle of discal cell. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis discussa Munro, 1935 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis ditissima (Munro, 1938)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pardalaspis) ditissima (Munro)
Pardalaspis ditissima Munro, 1938: 164.
Body length: 6.79 (6.0-7.5) mm; wing length: 6.79 (6.0-7.5) mm.

Male
Head. Antennal segments orange to dark red. Third antennal segment twice as long as second segment. Arista basal part orange, otherwise dark. Frons yellow in ground colour (in some specimens orange-red), silvery over entire length; with dispersed short hairs, same colour as frons. Ocellar triangle dark. Face dark red; no median band but lower margin white. Occiput moderately swollen below, pale whitish. Chaetotaxy normal for subgenus. Thorax. Ground colour of mesonotum greyish-brown, anterior border dark brown, sickle-shaped and extended along lateral margins; also with three poorly defined darker stripes. Postpronotum paler than mesonotum. Chaetotaxy normal for subgenus. Mesonotum with pale pilosity, brown part with dark hairs. Anepisternum along upper margin and lower half with darker hairs. Two anepisternal bristles, sometimes three, but then median one usually less well developed. Scutellum yellow with apical markings black; basally with two dark spots. Legs dark yellow to orange brown, femora with darker streaks; setation typical of subgenus. Hind femur with dorsal hairs dark apically. Setae general dark in colour, posterior row on front femur dark. Wings with brownish bands, yellow markings strongly reduced. Banding, setation and venation normal for subgenus. Marginal band continuous; discal band joined with marginal band; cross-vein r-m at middle of discal cell; vein R1 ending before cross-vein r-m. Abdomen. grey-brown, sometimes with orange tinge, with clearly defined spots. Pattern of spots and setation normal for subgenus.
Female
As male except for the following characters. Frons yellow, not silvery; with distinctly darker hairs. Face wholly yellow, lower margin same colour. Mesonotum without darker anterior part, concolorous; pilosity completely pale. Scutellum basally sometimes more whitish. Oviscape shorter than abdominal terga 3-6, orange in colour.
(Description after De Meyer, 1996)
See description of Ceratitis ditissima (Munro, 1938) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis divaricata (Munro, 1933)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) divaricata (Munro)
Hoplolophomyia divaricata Munro, 1933a: 31.
Body length: 6.32 (5.65-7.10) mm; wing length: 6.54 (6.15-6.85) mm.

Male
Head. Antenna yellow. Third antennal segment twice as long as second segment. Arista almost bare, extremely short hairs which are hardly visible. Frons with short scattered hairs which are usually distinctly darker than frons, sometimes largely the same colour; convex to flat, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum white or dirty white, unspotted. Ground colour of mesonotum pale with yellow-orange tinge; mesonotum, intra-alar spots sometimes less developed. Chaetotaxy normal for subgenus. Scapular setae pale. Two anepisternal bristles, lower bristle sometimes weaker and brownish coloured. Scutellum white basally, otherwise yellow with three black separate markings apically; basally without dark spots, but two subapical spots. Subscutellum pale with a single median spot. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale; ventral row yellow. Wing bands with extensively yellow markings. Banding, two longitudinal bands, marginal band continuous, band along vein M; cubital and medial bands not present; crossvein r-m before middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Arista with slightly longer hairs, but still very short and hardly visible. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis divaricata (Munro, 1933) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis dumeti Munro, 1933"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) dumeti Munro
Ceratitis (Pardalaspis) dumeti Munro, 1933a: 35.
Body length: 4.74 (4.15-5.10) mm; wing length: 4.29 (4.15-4.60) mm.

Male
Head. Antenna yellow-orange. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons with short scattered hairs which are distinctly darker than frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum yellow-white, paler than centre mesonotum; unspotted. Ground colour of mesonotum pale with yellow-orange tinge; mesonotum, sometimes prescutellar spots weakly connected with dorsocentral setae, presutural spots variable and occasionally median line present. Chaetotaxy normal for subgenus. Scapular setae pale. One or two anepisternal bristles, lower bristle usually smaller and less developed, occasionally palish colour. Scutellum white basally, otherwise yellow with three black separate markings apically; basally without dark spots, only slight darker colouration. Subscutellum three black spots at same position as apical scutellar spots, widely separated. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Ventral, posterior and posterodorsal rows on fore femur pale. Wing bands with extensively yellow markings. Banding, marginal band continuous; cubital band free, medial band absent; crossvein r-m before middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Antenna more yellow. Subscutellar markings narrowly separated (as in other Ceratalaspis spp.). Mesonotum with more outspoken orange tinge. Oviscape longer than abdominal terga 3-6 combined, or at least as long as terga.
(Description after De Meyer, 1998)
See description of Ceratitis dumeti Munro, 1933 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis edwardsi (Munro, 1957)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pardalaspis) edwardsi (Munro)
Pardalaspis edwardsi Munro, 1957: 868.
Body length: 8.0 (7.5-8.5) mm; wing length: 8.08 (7.5-9.0) mm.

Male
Head. Antennal segments yellow-orange. Third antennal segment three times as long as second segment. Arista dark over entire length. Frons yellow in ground colour,with indistinct silvery sheen especially in centre; with dispersed short hairs, distinctly darker than frons. Ocellar triangle dark. Face orange to dark orange; no median band. Occiput moderately swollen below, pale whitish. Chaetotaxy normal for subgenus. Thorax. Ground colour of mesonotum brownish; with three darker longitudinal stripes, central line broad, lateral ones interrupted and thinner, darker markings around prescutellar acrostichal setae. Postpronotum same colour as mesonotum. Chaetotaxy normal for subgenus. Mesonotum with mainly dark pilosity. Anepisternum along upper margin and lower half with darker hairs. Two anepisternal bristles. Scutellum yellow with apical markings black; basally with two dark spots. Legs yellow to orange-brown, setation typical of subgenus; predominantly dark. Posterior row on front femur dark. Wings with brownish bands, yellow markings strongly reduced. Banding, setation and venation normal for subgenus. Marginal band continuous; discal band joined with marginal band; cross-vein r-m at middle of discal cell; vein R1 ending before cross-vein r-m. Abdomen. Pale greyish, with clearly defined spots. Pattern of spots and setation normal for subgenus.
Female
As male except for the following characters. Face and frons variable, from pale yellow to yellow-orange. Mesonotum with pale pilosity except along longitudinal stripes where there are dark hairs. Cross-vein r-m before middle of discal cell; R1 ending before or opposite cross-vein r-m. Abdomen with orange tinge. Oviscape moderately long, at least as long as abdominal terga 3-6; orange in colour.
(Description after De Meyer, 1996)
See description of Ceratitis edwardsi (Munro, 1957) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis epixantha (Hering, 1941)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) epixantha (Hering)
Pardalaspis epixantha Hering, 1941: 69.
Body length: 4.00 mm; wing length: 4.00 mm.

Male
Head. Antenna yellow. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons with short scattered hairs which are largely the same colour as frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum pale yellow, unspotted. Ground colour of mesonotum with dark orange tinge. Chaetotaxy normal for subgenus. Scapular setae, one outer pale, others missing. Two anepisternal bristles, lower one less well developed. Scutellum white basally, otherwise yellow with three black separate markings apically; basally without dark spots, only slight darker colouration. Subscutellum with three separate spots at same position as apical scutellar spots. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Ventral spines pale. Wing bands with yellow-brown markings. Banding; marginal band continuous; cubital band free, medial band absent; crossvein r-m well before middle of discal cell, at basal two-fifth. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Dark yellow. Setation and banding typical for subgenus.
Female
Unknown.
See description of Ceratitis epixantha (Hering, 1941) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis fasciventris (Bezzi, 1920)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) fasciventris (Bezzi)
Pterandrus rosa var. fasciventris Bezzi, 1920: 228.
Body length: 4.47 (3.95-5.15) mm; wing length: 4.96 (4.45-5.75) mm.

Male
Head:
Antenna yellowish orange. First flagellomere 2-3 times as long as pedicel. Arista with short to moderately long rays; ventral rays shorter and sparser than dorsal rays, especially basally. Frons yellow; with short scattered setulae distinctly darker than frons. Frontal setae well developed. Face yellowish white. Genal seta and setulae dark, well developed. Thorax:
Postpronotal lobe yellowish white, without spot. Mesonotum:
ground color dark gray, sometimes with orange tinge; with streaks and darker markings but without distinct spots, except prescutellar white separate markings, usually with paler gray area in between, occasionally merged. Scapular setae dark. Scutellum yellowish white, basally usually without dark spots, rarely with two separate spots; apically with three separate dark spots, extending to basal half, sometimes to basal 0.33. Anepisternum on ventral half yellowish brown; setulae pale. Legs:
Yellow except where otherwise noted; setation typical for subgenus, mixed pale and dark. Foreleg:
at most slightly darker yellow anteriorly; femur without bushy feathering posteriorly, only row of dispersed, long and usually black setulae posteriorly, setulae shorter and pale posterodorsally and posteroventrally; ventral setae black. Midleg:
femur with dispersed pale setulae at base ventrally; tibia not broadened; anteriorly with inconspicuous silvery shine on distal half when viewed from certain angle; ventrally and dorsally with dark feathering along distal half. Hindleg:
femur at apical 0.25 with longer setulae dorsally and ventrally. Wing:
bands brown or yellowish brown. Interruption between marginal and discal bands near vein R1 clear and complete; cubital band free; medial band absent; crossvein R-M opposite middle of discal cell, sometimes just proximal to middle. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu oblique anterobasally. Abdomen:
Mostly yellow. Tergites 2 and 4 with pale gray band on posterior half, anterior margin sometimes with narrowly brownish colored. Tergite 3 with distinct brownish black transverse band along posterior half; rarely more complete brown. Tergite 5 with basal half brownish, sometimes divided medially into two spots, or only narrowly brownish along anterior margin.
Male epandrium in lateral view with posterior lobe of lateral surstylus short and straight, anterior lobe well pronounced.
Female
As male except for the following characters:
Anepisternum on ventral half brown or yellowish brown. Crossvein DM-Cu variable. Legs without feathering; forefemur with posterodorsal row usually partly dark; forefemur posteroventrally with pale pilosity. Oviscape shorter than preabdomen. Aculeus at most six times longer than wide; tip with distinct apical indentation and lateral margin slightly sinuous.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis fasciventris (Bezzi, 1920) in source PDF."""
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Marc de Meyer

"""Ceratitis flexuosa (Walker, 1853)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) flexuosa (Walker)
Trypeta flexuosus Walker, 1853: 382.
Body length: 4.52 (3.75-4.75) mm; wing length: 5.00 (3.85-5.45) mm.

Male
Head:
Antenna yellowish orange. First flagellomere twice as long as pedicel. Arista with short to moderately long rays; ventral rays shorter and sparser than dorsal rays, especially basally. Frons convex, pale yellow; with short scattered setulae largely of same color as frons. Frontal setae well developed. Face yellowish white; gena somewhat darker. Genal seta and setulae dark, well developed. Thorax:
Postpronotal lobe yellowish white, without spot. Mesonotum:
ground color pale gray with faint orange tinge; microtrichose silvery shine; with streaks and darker markings but without distinct spots, except at transverse suture and around dorsocentral and (to lesser extent) prescutellar seta implant dark, shiny spots, prescutellar yellowish white markings separate. Scapular setae dark. Scutellum yellowish white, basally without spots, apically with three strongly reduced dark spots. Subscutellum yellowish orange, without distinct spots. Anepisternum on ventral half darker yellow but often not pronounced; setulae pale. Legs:
Yellow except where otherwise noted; setation typical for subgenus, mixed pale and dark. Foreleg:
femur without bushy pilosity, with dispersed dark setulae, ventral setae black. Midleg:
femur anteriorly with large median white spot; along dorsal edge on distal half to 0.6 with short dense black pilosity, ventrally with few long black setulae near base; tibia dilated, with apical 0.33 to 0.4 brown with silvery spot, only conspicuous when viewed from certain angle; with black feathering dorsally and ventrally along distal 0.4 to half; anteriorly with third row of very short black feathering, area between third and dorsal row not dark. Hindleg:
femur at apical 0.25 with longer setulae dorsally and ventrally. Wing:
bands largely reduced to isolated brownish spots, especially marginal band. Medial band absent; crossvein R-M just proximal to middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu oblique posterobasally. Abdomen:
Mostly yellowish orange. Tergites 2, 4 and 5 largely grayish (not silvery); tergite 3 with posterior 0.33 brownish, interrupted medially; tergite 5 with anterior part brownish or yellowish brown, usually as two separate spots, occasionally as continuous band.
Male epandrium in lateral view with posterior lobe of lateral surstylus short and stout, slightly curved; anterior lobe well pronounced.
Female
As male except for the following characters:
Darker patches on gena not distinct. Subscutellum usually with obscured dark spot in the middle. Legs without feathering; forefemur posteroventrally with pale pilosity. Abdominal tergite 5 anteriorly at most with darker yellow markings. Oviscape shorter than preabdomen. Aculeus at most six times longer than wide; tip with distinct apical indentation and lateral margin slightly sinuous. Some female specimens have a generally darker appearance.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis flexuosa (Walker, 1853) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis fulicoides (Munro, 1943)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) fulicoides (Munro)
Pterandrus fulicoides Munro, 1943: 137.
Body length: 5.84 (5.60-6.25) mm; wing length: 6.51 (5.80-7.00) mm.

Male
Head:
Antenna yellowish orange. First flagellomere 2-3 times as long as pedicel. Arista with short to moderately long rays; ventral rays shorter and sparser than dorsal rays, especially basally. Frons convex, with poorly developed median protuberance; pale, center yellow; with short scattered setulae largely of same color as frons. Frontal setae absent; posterior orbital less developed. Face yellowish white. Genal seta pale or reddish; genal setulae dark reddish, poorly developed. Thorax:
Postpronotal lobe yellowish white, without spot. Mesonotum:
ground color pale gray with distinct orange tinge, microtrichose silvery shine; with streaks and darker markings but without distinct spots, except at transverse suture and around dorsocentral and prescutellar setae, prescutellar yellow-white markings separate. Scapular setae dark. Scutellum yellowish white, basally without distinct spots, apically with three separate black spots, extending to basal half. Anepisternum on ventral half yellowish orange; setulae pale. Legs:
Yellow except where otherwise noted; setation typical for subgenus, mainly pale. Foreleg:
femur posteriorly with bush of long orange setulae along entire length, posterodorsal setulae longer; ventrally with row of long orange setae. Midleg:
femur ventrally with row of long orange setulae, on distal 0.33 closely appressed to form feathering; midtibia with orange feathering dorsally along distal 0.66 and ventrally along distal 0.8; anteriorly with short to moderately long dispersed orange setulae, longer along dorsal row. Hindleg:
femur at apical 0.25 with longer setulae dorsally and to a lesser extent ventrally. Wing:
bands brownish or yellowish brown. Marginal band only well developed on distal half, interruption between marginal and discal bands broad and complete; discal band narrow and partly interrupted near crossvein R-M; cubital band free; medial band absent; crossvein R-M slightly beyond middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu oblique posterobasally. Abdomen:
Mostly yellow. Tergite 2 largely and tergite 4 across posterior half with pale gray (not silvery) band; tergite 3 across posterior 0.33 to 0.25 with brown band, interrupted medially or with median notch; tergite 5 across posterior margin sometimes narrowly brownish.
Male epandrium in lateral view with lateral surstylus slender and elongate; posterior lobe straight and elongate, anterior lobe well pronounced.
Female
As male except for the following characters:
Two (sometimes three) well developed frontal setae present; orbital setae stronger developed. Genal seta dark, genal setulae all dark and well developed. Wing bands slightly paler; discal and cubital bands broader and well developed; medial band weakly developed but present, joining marginal band. Legs without feathering. Forefemur with posterior and posterodorsal rows of dark orange setae; posteroventral setulae pale. Ventral setae orange, basally black. Abdominal tergite 5 not darkened posteriorly. Oviscape almost as long as preabdomen. Aculeus about eight times longer than wide; tip pointed and lateral margin slightly concave.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis fulicoides (Munro, 1943) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis grahami Munro, 1935"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) grahami Munro
Ceratitis (Pardalaspis) grahami Munro, 1935: 303.
Body length: 5.00 mm; wing length: 5.40 mm.

Male
Head. Antenna dark orange. Antennal inplant with darker horizontal band (but not very clear). Third antennal segment twice as long as second segment. Arista with hairs of moderate length over entire length (but not plumose). Frons with short scattered hairs which are largely the same colour as frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus except only one frontal bristle. Thorax. Postpronotum white, with small black spot. Ground colour of mesonotum pale with orange tinge. Chaetotaxy normal for subgenus. Scapular setae pale. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically; basally without dark spots, only slight darker colouration. Subscutellum completely dark. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Fore femur with dense orange setation ventrally and posterodorsally along entire length, not outspoken feathery but more bushy; similar setation on dorsal and ventral side along apical fourth of hind femur. Wing bands with markings extensively yellow. Banding, marginal band continuous; cubital band free and medial band absent; crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen. Brownish colour. Setation and banding typical for subgenus.
Female
Unknown.
(Description after De Meyer, 1998)
See description of Ceratitis grahami Munro, 1935 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis guttiformis Munro, 1935"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) guttiformis Munro
Ceratitis (Pardalaspis) guttiformis Munro, 1935: 309.
Body length: 5.80 (5.75-5.85) mm; wing length: 5.03 (4.65-5.40) mm.

Female
Head. Antenna yellow-orange. Third antennal segment three times as long as second segment. Arista with longer hairs but not distinctly plumose. Frons with short scattered hairs which are distinctly darker than frons (in paratype only slightly darker however); concave to flat, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum concolorous with ground colour of mesonotum, with black spot. Ground colour of mesonotum pale with orange tinge. Chaetotaxy normal for subgenus. Scapular setae black. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically; basally with 2 dark spots (poorly developed in holotype). Subscutellum pale with three dark separate spots. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing bands with markings extensively yellow. Banding, marginal band continuous; cubital band free or very weakly connected with discal band; medial band absent; crossvein r-m at or before middle of discal cell. Crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen. Yellow-orange colour. Setation and banding typical for subgenus. Oviscape almost as long as abdominal terga 3-6 combined.
Male
Unknown.
(Description after De Meyer, 1998)
See description of Ceratitis guttiformis Munro, 1935 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis hamata De Meyer, 1996"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pardalaspis) hamata De Meyer
Ceratitis (Pardalaspis) hamata De Meyer, 1996 : 21.
Body length: 6.57 (6.0-7.0) mm; wing length: 6.43 (6.0-7.0) mm.

Male
Head. Antennal segments orange. Third antennal segment twice as long as second segment. Arista basal part orange coloured, otherwise dark. Frons yellowish in ground color (in specimens from Yangambi more orange), silvery over entire length; with dispersed short hairs, at most slightly darker than frons. Ocellar triangle dark. Face orange to orange-red; no median band. Occiput moderately swollen below, pale whitish. Chaetotaxy normal for subgenus. Thorax. Ground colour of mesonotum greyish with orange tinge; also with three poorly defined darker stripes, usually interrupted; sometimes with darker spots along prescutellar acrostichal setae. Postpronotum slightly paler than mesonotum, no orange tinge. Chaetotaxy normal for subgenus. Mesonotum with pale pilosity. Anepisternum along upper margin with darker hairs, otherwise pale pilosity. Two anepisternal bristles. Scutellum yellow with apical markings black; basally with two dark spots, these variable and not always distinct. Legs dark yellow to orange brown; setation typical of subgenus. Setae dark, front femur with posterior row dark. Wings with brownish bands, yellow markings strongly reduced. Banding, setation and venation normal for subgenus. Marginal band continuous; discal band joined with marginal band; cross-vein r-m before middle of discal cell; vein R1 ending opposite cross-vein r-m. Abdomen. greyish, with clearly defined spots. Pattern of spots and setation normal for subgenus.
Female
As male except for the following characters. Third antennal segment two to three times as long as second. Frons darker, more orange (as in males of Yangambi, cf above); silvery shine less conspicuous than in male, along setal sockets however pronounced; hairs distinctly darker than frons. Face wholly yellow. Vein R1 ending before or opposite cross-vein r-m. Oviscape shorter than abdominal terga 3-6, orange in colour.
(Description after De Meyer, 1996)
See description of Ceratitis hamata De Meyer, 1996 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis hancocki De Meyer, 1998"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) hancocki De Meyer
Ceratitis (Ceratalaspis) hancocki De Meyer, 1998: 279.
Body length: 4.64 (4.60-4.70) mm; wing length: 4.59 (4.40-4.85) mm.

Male
Head. Antenna yellow-orange. Third antennal segment slightly less than three times as long as second segment. Arista with short to medium long hairs over entire length. Frons yellow; with short scattered hairs which are distinctly darker than the frons; in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum yellow, almost same colour as centre of mesonotum; postpronotum spotted but spot small. Ground colour of mesonotum pale with yellow-orange tinge. Chaetotaxy normal for subgenus. Scapular setae black, inner reddish. One anepisternal bristle. Scutellum mainly yellow; with three black separate markings apically except at ventral margin where narrowly joining; basally two dark distinct spots. Subscutellum pale with three dark separate spots except along dorsal edge where touching. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Ventral spines on fore femur reddish. Wing bands with markings yellow-brown. Banding as in fig. 1b, marginal band continuous; cubital band joined with discal band; medial band absent; crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As male except for the following characters:
postpronotal spot sometimes obscured. Scapular setae and ventral spines darker. Mesonotum with postsutural median spot absent. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis hancocki De Meyer, 1998 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis lentigera Munro, 1933"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) lentigera Munro
Ceratitis (Pardalaspis) lentigera Munro, 1933b: 8.
Body length: 4.51 (3.85-5.40) mm; wing length: 4.28 (3.75-4.85) mm.

Male
Head. Antenna yellow-orange. Third antennal segment three times as long as second segment. Arista with short to medium length hairs over entire length. Antennal inplant with dark horizontal band. Frons yellow with dark median band; with short scattered hairs which are largely the same colour as frons; convex but not distinct, in lateral view not distinctly projecting forwards at antennal implant. Face yellow-white with dark median band; lower eye-margin with slightly darker colouration, not distinct patch. Chaetotaxy normal for subgenus. Bands on face and frons not always well pronounced. Thorax. Postpronotum white, spotted. Ground colour of mesonotum pale with orange tinge; mesonotum, colouration along median line variable. Chaetotaxy normal for subgenus. Outer scapular setae black, inner ones pale (sometimes all black). One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically; with two separate dark spots placed subbassaly, hardly touching base of scutellum. Subscutellum with three black spots at same position as scutellar spots, dorsally narrowly joining. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing bands with extensively yellow markings, sometimes more yellow-brown. Banding, marginal band continuous; cubital band free, medial band absent; crossvein r-m just before middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Pale yellow. with few spots, no distinct banding.
Female
As male except for the following characters:
Facial and frontal bands sometimes less clear, and interrupted. Mesonotum, orange tinge sometimes very faint. Abdomen greyish brown. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis lentigera Munro, 1933 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis lepida (Munro, 1969)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) lepida (Munro)
Pterandrus lepidus Munro, 1969: 420.
Body length: 4.81 mm; wing length: 5.25 mm.

Male
Head:
Antenna orange-brown. First flagellomere three times as long as pedicel. Arista missing in holotype. Frons flat, yellow; with short scattered setulae distinctly darker than frons. Frontal setae missing in holotype (probably broken). Face yellowish white; gena dark, darkening partly continued along ventral margin but not in median part; mouthparts black forming a contrasting character together with dark gena when viewed anteriorly. Genal seta black; genal setulae black, well developed. Thorax:
Postpronotal lobe white, without spot. Scutal pattern:
ground color dark gray with silvery shine; with streaks and darker markings but without distinct spots except prescutellar yellow separate markings. Scapular setae dark. Scutellum yellowish white, basally without spots, apically with three separate dark spots extending to halfway scutellar disc. Anepisternum on ventral half black; setulae on ventral half and across posterior margin black, otherwise pale. Legs:
Yellow except where otherwise noted; setation typical for subgenus, mainly dark. Foreleg:
coxa black; femur anteriorly with contrasting black/white pattern, white spot and basal part of femur turn silverly when viewed from certain angle, anteroventrally with bush of short dark setulae along medial half; posteriorly dark except apically along ventral side; with dispersed bush of long dark setulae along entire length, posterodorsal setulae longer; ventral setae dark; tibia anteriorly mainly brownish black, only distally and along ventral side partly yellowish. Midleg:
femur anteriorly largely brownish black, only distal end and along dorsal margin partly yellowish; ventrally with dark feathering along entire length except for small interruption (two-three setae wide) in median part; tibia broadened and anteriorly largely brownish black, weak silvery when viewed from certain angle; with black feathering dorsally along distal 0.9 and ventrally along distal 0.66, second row of short black feathering present anterior to dorsal feathering, closely apressed to first one (not easy to differentiate); anteriorly with dispersed short black setulae. Hindleg:
femur brown except apically; at apical 0.25 with longer setulae dorsally and ventrally. Wing:
bands yellow and brown. Interruption between marginal and discal bands near vein R1 complete; cubital band free; medial band absent; crossvein R-M just proximal to middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu straight. Extension cup fairly longer. Abdomen:
Mostly orange-brown. Tergite 1 dark brown across posterior margin. Tergites 2 and 4 with silvery band occupying most if not whole of tergite. Tergite 3 dark brown posteriorly. Tergite 5 with weak silvery transverse band along posterior margin.
Male terminalia not dissected.
Female
Unknown.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis lepida (Munro, 1969) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis lineata (Hering, 1938)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) lineata (Hering)
Pardalaspis lineata Hering, 1938: 406.
Body length: 4.60 mm; wing length: 4.50 mm.

Male
(Based on original description, with additions based on KBIN and USNM specimens and indicated as such) Head. Antenna yellow to yellow-orange. Third antennal segment broken in holotype, less than three times as long as second segment in USNM specimen. Arista with short hairs over entire length. Frons (in material studied) yellow, with scattered hairs which are distinctly darker than frons, in lateral view not distinctly projecting forwards at antennal implant; face yellow-white, lower margin near mouth and below eyes more yellow. Chaetotaxy normal for subgenus. Thorax. Postpronotum yellow-white, spotted. Ground colour of mesonotum pale with orange tinge. Inner scapular setae missing, outer scapular setae black. One anepisternal bristle, in USNM specimen on one side second lower bristle which is less developed. Scutellum white basally, otherwise yellow with three black separate markings apically; basally two dark separate spots. Subscutellum with three black spots, widely separated, dorsally connected. Wing bands with markings extensively yellow. Banding, marginal band continuous; cubital band free; medial band absent; crossvein r-m at or just before middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Ventral spines dark in USNM specimen. Abdomen yellow. Setation and banding typical for subgenus.
Female
Unknown.
(Description after De Meyer, 1998)
See description of Ceratitis lineata (Hering, 1938) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis lunata Munro, 1935"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) lunata Munro
Ceratitis (Pardalaspis) lunata Munro, 1935: 305.
Body length: 4.49 (3.85-5.00) mm; wing length: 4.65 (4.40-4.90) mm.

Male
Head. Antenna yellow-orange. Third antennal segment slightly more than twice as long as second segment. Arista distinctly plumose. Frons with short scattered hairs which are largely the same colour as frons, sometimes slightly darker; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum white, with relatively small black spot. Ground colour of mesonotum pale with orange tinge; mesonotal pattern, prescutellar spots not touching intra-alar spots, median sutural spot missing. Chaetotaxy normal for subgenus. Inner scapular setae pale, outer ones black. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically; basally with two dark spots. Subscutellum pale with three dark separate spots, except along dorsal margin where touching. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing bands with markings extensively yellow. Banding, marginal band continuous; cubital band free; medial band absent; crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. More or less brownish. Setation and banding typical for subgenus.
Female. As male except for the following characters:
arista with slightly shorter hairs. Postpronotal spot larger. Outer scapular seta (only present in paralectotype) reddish (inner missing in all specimens). Subscutellum completely dark. Wing with crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen yellow in paratype. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis lunata Munro, 1935 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis malgassa Munro, 1939"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratitis) malgassa Munro
Ceratitis (Ceratitis) malgassa Munro, 1939: 141.
Body length: 5.06 (4.45-5.55) mm; wing length: 5.01 (4.65-5.55) mm.

Male
Head. Antenna yellow or yellow-orange. Third antennal segment twice as long as second segment. Arista almost bare, at most with very short hairs on base. Frons yellow with orange patches; with short scattered hairs which are largely the same colour as frons. Frontal and orbital bristles pale, upper orbital sometimes dark, ocellar black; lower orbital modified, about as long as arista, with apical end white and narrowly spatulate shaped, upper orbital and frontals partly reduced, only moderately developed. Face yellow-white, lower eye margin darker yellow. Genal bristle pale; genal setulae pale, weakly developed. Postocellar and outer vertical pale. Thorax. Postpronotum yellow-white, with distinct black spot. Mesonotal pattern:
ground colour orange-brown, microtrichiae pattern silvery with brownish shine, spots black-brown except rather indistinct sutural yellow or yellow-white spots, prescutellar yellow markings merged. Scapular setae pale. Scutellum yellow-white, basally with two dark narrowly merged spots, apically with three merged spots, only slightly incised. Anepisternum with lower half darker yellow but not distinct, hairs in lower half darker yellow. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Fore femur posteriorly with bush of longer orange-coloured hairs along entire length, posterodorsal hairs longer. Anteroventrally row of longer hairs, as long as posterodorsal hairs; ventral spines yellow-orange. Mid femur posteroventrally with row of longer hairs. Hind femur with longer hairs dorsally and ventrally on apical fourth. Wing. Banding; marginal band with narrow interruption, sometimes partly touching; cubital band free; medial band joined with marginal band; crossvein r-m just beyond middle of discal cell. Vein R  before crossvein r-m. Orientation crossvein dm-cu variable. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As in male except for the following characters:
Arista with slightly longer hairs. Hairs on frons distinctly darker than frons. Darker colouration on lower eye margin not outspoken. Frontal, orbital and ocellar bristles black, orbitals not modified, frontal bristles well developed. Face sometimes darker. Genal bristles dark or pale; genal setulae dark, strongly developed. Incisions of apical scutellar spots not as deep as in male. Legs without feathering; ventral spines on fore femur black. Oviscape shorter than abdominal terga.
(Description after De Meyer, 2000)
See description of Ceratitis malgassa Munro, 1939 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis manjakatompo Hancock, 1984"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratitis) manjakatompo Hancock
Ceratitis (Ceratitis) manjakatompo Hancock, 1984: 283.
Body length: 3.54 (3.5-3.6) mm; wing length: 3.6 (3.55-3.65) mm.

Female
Head. Antenna yellow-orange. Third antennal segment three times as long as second segment. Arista with short hairs over entire length, longer than in other Ceratitis species (except C. cornuta). Frons pale yellow to yellow; with short scattered hairs which are largely the same colour as frons, at most slightly darker. All cephalic bristles dark, except postocellar and outer verticals. Face yellow-white. Genal bristles dark reddish, strongly developed. Thorax. Postpronotum, white with large black spot. Mesonotal pattern:
ground colour orange-brown, microtrichiae pattern silvery with brownish shine, spots black-brown except conspicuous sutural white spots, prescutellar white markings merged. Scapular setae pale. Scutellum yellow-white, basally with two dark separate spots, apically with three merged spots, weakly incised. Basal spots weakly joined with apical spots. Anepisternum with lower half darker yellow but not distinct. Legs yellow; setation typical for subgenus, mainly yellow or reddish. Ventral spines on fore femur reddish. Wing bands with markings yellow. Banding; marginal band with clear and complete interruption; cubital band free; medial band narrow, joined with marginal; crossvein r-m beyond middle of discal cell. Vein R  before or equal with crossvein r-m. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus. Oviscape shorter than abdominal terga.
Male
Unknown.
(Description after De Meyer, 2000)
See description of Ceratitis manjakatompo Hancock, 1984 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis marriotti Munro, 1933"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) marriotti Munro
Ceratitis (Pardalaspis) marriotti Munro, 1933a: 36.
Body length: 5.12 (4.60-5.25) mm; wing length: 5.02 (4.75-5.25) mm.

Male
Head. Antenna yellow. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons with short scattered hairs which are usually darker than frons, sometimes paler and almost same colour as frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Lower eye margin with darker marking (not always very distinct). Chaetotaxy normal for subgenus. Thorax. Postpronotum white, unspotted. Ground colour of mesonotum pale with orange tinge; mesonotal pattern, median spot between postpronota variable, sometimes only present as a darker yellow patch. Chaetotaxy normal for subgenus. Scapular setae black. One anepisternal bristle. Scutellum white with three black separate markings apically, these markings hardly visible in dorsal view, restricted to apical margin; basally without dark spots, only slight darker colouration. Subscutellum completely dark. Legs yellow; setation typical for subgenus, mixed darkish and pale. Posterior and posterodorsal rows pale, sometimes dark posteriorly. Wing bands with yellow-brown markings, sometimes more extensively yellow. Banding, marginal band with clear interruption after cell R1; cubital band free; medial band joined with marginal band; crossvein r-m at or before middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow (sometimes brownish discolored). Setation and banding typical for subgenus.
Female
As male except for the following characters:
Frons with scattered hairs that are always darker than frons. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis marriotti Munro, 1933 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis melanopus (Hering, 1942)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) melanopus (Hering)
Pardalaspis melanopus Hering, 1942: 282.
Body length: male 3.35, female 5.40 mm; wing length: male 4.20, female 5.60
mm.

Male
Head:
Antenna brownish. First flagellomere twice as long as pedicel. Arista with short rays. Frons convex; yellowish orange with weak silvery shine; with short scattered setulae of same color as frons. Frontal setae well developed. Face yellowish orange. Thorax:
Postpronotal lobe white, not spotted. Scutal pattern, ground color darkish gray to brown, microtrichose areas with ashgray shine, poorly developed lateral yellow lines, prescutellar white markings separate, with grayish area in between. Scapular setae dark. Scutellum white, basally without dark spots, apically more yellowish with three separate spots. Legs:
orange to brown; setation typical for subgenus, dark. Without feathering. Wing:
Bands brown. Interruption between marginal and discal bands near vein R clear and complete; cubital band free; medial band absent; crossvein R-M proximal to middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu oblique posterobasally. Abdomen:
Mostly pale orange. Tergite 3 with posterior half brown. Setation and banding typical for subgenus.
Male terminalia not dissected.
Female
As male except for the following characters:
Rays on arista longer but not plumose. First flagellomere longer. Face yellow. Scutal pattern slightly different, with more obscured orange lateral stripes, area between prescutellar markings more orange. Marginal band partly interrupted. Oviscape about as long as preabdomen. Aculeus at least eight times longer than wide; tip with distinct apical indentation and lateral margin more or less straight.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis melanopus (Hering, 1942) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis mlimaensis De Meyer, 1998"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) mlimaensis De Meyer
Ceratitis (Ceratalaspis) mlimaensis De Meyer, 1998: 281.
Body length: 4.31 (4.25-4.45) mm; wing length: 4.54 (4.40-4.70) mm.

Male
Head. Antenna yellow-orange. Third antennal segment two times as long as second segment. Arista with short hairs over entire length. Frons yellow to orange; with short scattered hairs which are largely the same colour as the frons; flat to convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum only slightly paler in colour than centre of mesonotum, yellow-white; unspotted. Ground colour of mesonotum pale with yellow-orange tinge; presutural median spot absent (in BMNH specimen, invisible in USNM specimen because of insect pin). Chaetotaxy normal for subgenus. Scapular setae black. One or two anepisternal bristles, if second present then lower reduced in size. Scutellum mainly yellow; with three black separate markings apically except at ventral margin where narrowly joining; basally no distinct spots. Subscutellum pale with three dark separate spots. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing bands with markings yellow-brown. Banding, marginal band continuous; cubital band free; medial band joined with marginal band; crossvein r-m before middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Mesonotum with presutural median spot present. Crossvein r-m well before middle of discal cell. Oviscape as long as abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis mlimaensis De Meyer, 1998 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis morstatti Bezzi, 1912"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) morstatti Bezzi
Ceratitis morstatti Bezzi, 1912: 12.
Body length: 5.13 (4.50-5.70) mm; wing length: 5.34 (5.00-5.80) mm.

Male
Head:
Antenna yellow-orange (yellow according to original description). First flagellomere twice as long as pedicel. Arista with medium long rays. Frons convex, yellow (dark red, with gray basal stripes according to orginal description); with short scattered setulae distinctly darker than frons. Genal seta black; genal setulae black, well developed. Thorax:
Postpronotal lobe white, unspotted. Scutal pattern, ground color pale gray (occasionally darker gray) with silvery shine; with very light streaks and darker markings but without distinct spots except prescutellar yellow separate markings. Scapular setae dark. Scutellum grayish, basally without spots, apically with three separate black spots extending to halfway scutellar disc. Anepisternum on ventral margin yellowish brown; setulae white, along ventral margin black. Legs:
Yellow; setation typical for subgenus, mainly dark; without feathering. Wing:
Bands yellow and brown. Interruption between marginal and discal bands near vein R1 complete; cubital band free (only occasionally weakly connected); medial band absent; crossvein R-M opposite middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu variable. Abdomen:
Mostly gray. Tergites 2 and 4 with pale gray band occupying most if not whole of tergite. Tergite 3 dark brown posteriorly. Tergite 5 with yellowish brown and brown patches.
Male terminalia not dissected.
Female
As male except for the following characters. Oviscape as long as preabdomen. Aculeus at least eight times longer than wide; tip with distinct apical indentation and lateral margin slightly sinuous.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis morstatti Bezzi, 1912 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis munroi De Meyer, 1996"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pardalaspis) munroi De Meyer
Ceratitis (Pardalaspis) munroi De Meyer, 1996 : 22.
Body length: 7.0 mm; wing length: 7.0 mm.

Female
Head. Antennal segments orange. Third antennal segment twice as long as second segment. Arista basal part yellowish, otherwise dark. Frons pale yellowish in ground color, weak silvery shine in lower part; with dispersed short hairs, distinctly darker than frons. Ocellar triangle dark. Face pale yellow. Occiput moderately swollen below, pale whitish. Chaetotaxy normal for subgenus. Thorax. Ground colour of mesonotum beige with conspicuous orange tinge; longitudinal lines hardly discernible. Postpronotum slightly paler than mesonotum. Chaetotaxy normal for subgenus. Mesonotum with pale pilosity. Anepisternum along upper margin and lower half with darker hairs. Two anepisternal bristles. Scutellum yellow with apical markings black; basally without dark spots, only slightly darker coloration. Legs yellow-orange, setation typical of subgenus. Setae mainly dark, front femur with posterior row dark. Wings with yellow-brown bands. Banding, setation and venation normal for subgenus. Marginal band continuous; discal band joined with marginal band; cross-vein r-m at middle of discal cell; vein R ending before cross-vein r-m. Abdomen. Pale with orange tinge, with clearly defined but small spots. Pattern of spots and setation normal for subgenus. Oviscape shorter than abdominal terga 3-6, yellow in colour
Male
Unknown.
See description of Ceratitis munroi De Meyer, 1996 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis nana Munro, 1933"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) nana Munro
Ceratitis (Pardalaspis) nana Munro, 1933b: 9.
Body length: 3.20 mm; wing length: 3.50 mm.

Male
Head. Antenna yellow. Third antennal segment three times as long as second segment. Arista with short to medium long hairs over entire length. Frons with short scattered hairs which are largely the same colour as frons; flat, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum white, spotted. Ground colour of mesonotum pale, with very faint orange tinge. Chaetotaxy normal for subgenus. Scapular setae pale. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically, median spot more or less diamond shaped; two dark separate spots subbassaly, hardly touching base of scutellum. Subscutellum with three dark spots, same position as scutellar spots. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal and ventral rows on fore femur pale. Wing bands with markings extensively yellow; cubital band free, medial band absent; crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow in ground colour. Weak spotted pattern, no distinct banding.
Female
Unknown.
(Description after De Meyer, 1998)
See description of Ceratitis nana Munro, 1933 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis neostictica De Meyer, 1998"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) neostictica De Meyer
Ceratitis (Ceratalaspis) neostictica De Meyer, 1998: 283.
Body length: 4.20 (4.00-4.40) mm; wing length: 4.37 (4.20-4.60) mm.

Male
Head. Antenna yellow-orange. Third antennal segment three times as long as second segment. Arista with medium long hairs over entire length. Frons yellow-orange, with short scattered hairs which are largely the same colour as frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum white, spotted. Ground colour of mesonotum pale with yellow-orange tinge. Chaetotaxy normal for subgenus. Scapular setae pale. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically; basally with two dark separate spots, only slight darker colouration. Subscutellum completely dark. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Fore femur posterodorsally with bushy pile of yellow-orange hairs; mid tibia and hind femur at apical third with feathering of same colour. Posterior and posterodorsal rows on fore femur pale. Ventral bristles on femora pale. Wing bands with extensively yellow markings. Banding, cubital band free, marginal band continuous; medial band free; crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Medial band not outspoken, at most weakly present. Legs without feathery or bushy pile. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis neostictica De Meyer, 1998 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis ovalis Munro, 1935"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) ovalis Munro
Ceratitis (Pardalaspis) ovalis Munro, 1935: 310.
Body length: 4.00 mm; wing length: 4.40 mm.

Male
Head. Antenna yellow-orange. Third antennal segment twice to three times as long as second segment. Arista distinctly plumose, with very long hairs over entire length. Frons with short scattered hairs which are largely the same colour as frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum white, with black spot. Ground colour of mesonotum pale with orange tinge. Chaetotaxy normal for subgenus. Scapular setae obscured in holotype. One anepisternal bristle. Scutellum yellow with three black separate markings apically; basally with two dark spots joining. Subscutellum completely dark. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing bands with extensive yellow markings. Banding, marginal band continuous; cubital band free; medial band absent; crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. cup extension only weakly sinoid. Abdomen. Orange-brown colour. Setation and banding typical for subgenus.
Female
Unknown.
(Description after De Meyer, 1998)
See description of Ceratitis ovalis Munro, 1935 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis paradumeti De Meyer, 1998"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) paradumeti De Meyer
Ceratitis (Ceratalaspis) paradumeti De Meyer, 1998 : 283.
Body length: 6.00 (5.90-6.10) mm; wing length: 5.30 (5.10-5.50) mm.

Female
Head. Antenna yellow-orange. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons with short scattered hairs which are largely the same colour as frons, at most slightly darker; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum pale yellow, unspotted. Ground colour of mesonotum pale with yellow-orange tinge. Chaetotaxy normal for subgenus. Scapular setae pale. Two anepisternal bristles. Scutellum white basally, otherwise yellow with three black separate markings apically; basally without dark spots, only slight darker colouration. Subscutellum three black spots at same position as apical scutellar spots. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Ventral, posterior and posterodorsal rows on fore femur pale. Ventral spines dark reddish. Wing bands with extensively yellow markings. Banding as in fig. 1a, marginal band continuous; cubital band free, medial band absent; crossvein r-m before middle of discal cell (in paratype at anterior third). Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus. Oviscape about twice as long as abdominal terga 3-6 combined.
Male
Unknown.
(Description after De Meyer, 1998)
See description of Ceratitis paradumeti De Meyer, 1998 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis penicillata Bigot, 1891"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) penicillata Bigot
Ceratitis ? penicillatus Bigot, 1891: 381.
Body length: 4.40 (3.90-5.15) mm; wing length: 4.78 (4.45-5.25) mm.

Male
Head:
Antenna yellowish orange, First flagellomere brown; twice or three times as long as pedicel. Arista with moderately long rays; ventral rays shorter and sparser than dorsal rays, especially basally. Frons yellow, sometimes pale yellow; darker patches on ventral half, especially at frontofacial angle; with short scattered setulae largely of same color as frons (occasionally darker). Frontal setae well developed. Face pale, median part with darker yellow to brown patch. Genal seta and setulae dark, well developed. Thorax:
Postpronotal lobe white, without spot. Scutal pattern:
ground color dark gray; with streaks and darker markings but without distinct spots except prescutellar white separate markings, with pale area in between. Scapular setae dark. Scutellum white, basally without spots, apically with three separate black spots, extending to basal half. Anepisternum along ventral margin only brown; setulae pale. Legs:
Yellow except where otherwise noted; setation typical for subgenus, mixed pale and dark. Foreleg:
femur with dorsal part brownish colored anteriorly and posteriorly; posteriorly no bush, only dispersed setulae; ventral setae black. Midleg:
femur anteriorly with large brown patch in median part, basally with few dispersed long pale setulae; tibia moderately broadened and anteriorly brownish black on distal half (except along dorsal row of feathering where yellow covered by short dark setulae), with black feathering ventrally along distal half, dorsally slightly over distal half. Hindleg:
femur brown on dorsal half; at apical 0.25 with longer setulae dorsally and ventrally. Wing:
bands yellowish brown to brown. Marginal band forming continuous band with anterior part of discal band; cubital band free; medial band absent (in rare cases a weak and incomplete band is present); crossvein R-M proximal to middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Orientation crossvein DM-Cu variable. Abdomen:
Mostly yellow. Tergites 2 and 4 across posterior half with pale gray band. Tergite 3 brown on posterior half. Tergites 4 and 5 along anterior margin yellowish brown, latter posteriorly narrowly brown.
Male epandrium in lateral view with lateral surstylus long and slender; posterior lobe straight and elongate.
Female
As male except for the following characters. First flagellomere darker but not brown. Face without darker coloration in median part. Crossvein DM-Cu oblique anterobasally. Medial band in one paratype of fumitactus weakly present. Legs without feathering, femora yellowish brown; forefemur posteroventrally with pale pilosity. Abdominal tergite 3 sometimes almost completely brown; tergite 5 on anterior 0.33 yellowish brown to brown, posteriorly without brownish margin. Oviscape slightly shorter than preabdomen. Aculeus at least eight times longer than wide; tip with distinct apical indentation and lateral margin weakly sinuous.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis penicillata Bigot, 1891 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis pinax Munro, 1933"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratitis) pinax Munro
Ceratitis (Pinacochaeta) pinax Munro, 1933: 34.
Body length: 3.96 (3.65-4.45) mm; wing length: 4.11 (3.90-4.45) mm.

Male
Head. Antenna yellow. Third antennal segment twice as long as second segment. Arista with short hairs. Frons yellow, sometimes with two whitish but indistinct spots near antennal implant; with short scattered hairs which are largely the same colour as frons. Frontal, ocellar and orbital bristles black; lower frontal weakly developed; lower orbital modified, ribbon shaped, black and slightly shorter than arista, tip variable usually blunt, sometimes pointed, rarely flattened and slightly widened. Face yellow-white, lower eye margin with slightly darker colouration. Genal bristle and setulae reddish, strongly developed. Postocellar and outer vertical pale. Thorax. Postpronotum white, spot more orange-brown, not black, sometimes indistinct. Mesonotal pattern:
ground colour orange-brown, microtrichiae pattern silvery with brownish shine, spots brown-black, sutural yellow-white spots present but sometimes indistinct, prescutellar yellow-white markings merged. Scapular setae pale. Scutellum yellow-white, basally with two dark separate spots, apically with three merged spots, usually incised to halfway. Anepisternum with lower half darker yellow, dark pilosity in lower half. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Fore femur posteriorly with bush of longer dark reddish hairs along entire length, basally usually darker and almost black; posterodorsal hairs longer; short anteroventral hairs and ventral spines black. Hind femur with longer hairs dorsally and ventrally on apical fourth. Wing. Banding; marginal continuous, sometimes with incomplete interruption; cubital band usually free, rarely joined with discal band; medial band absent; crossvein r-m at middle of discal cell. Vein R  before or equal with crossvein r-m. Crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As in male except for the following characters:
Antenna yellow-orange; third antennal segment two to three times longer than second segment. Orbitals not modified. Both frontal bristles well developed. Genal bristles dark. Anepisternum without dark pilosity. Crossvein r-m at or just beyond middle of discal cell. Crosvein dm-cu posteriorly more inwards than anteriorly. Vein R  equal with crossvein r-m. Oviscape shorter than abdominal terga.
(Description after De Meyer & Freidberg, in press)
See description of Ceratitis pinax Munro, 1933 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis punctata (Wiedemann, 1824)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pardalaspis) punctata (Wiedemann)
Tephritis punctata Wiedemann, 1824: 55.
Body length: 6.86 (6.0-7.5) mm; wing length: 6.86 (6.5-7.0) mm.

Male
Head. Antennal segments orange-red. Third antennal segment twice as long as second segment. Arista basal part orange coloured, otherwise dark. Frons yellowish in ground colour, silvery over entire length; with dispersed short hairs, distinctly darker than frons. Ocellar triangle dark. Face orange to orange-red; no median band. Occiput moderately swollen below, pale whitish. Chaetotaxy normal for subgenus. Thorax. Ground colour of mesonotum greyish to grey-brown; also with three poorly defined darker stripes, usually interrupted; sometimes with darker spots along prescutellar acrostichal setae. Postpronotum same colour as mesonotum. Chaetotaxy normal for subgenus. Mesonotum with pale pilosity. Anepisternum along upper margin with darker hairs, otherwise pale pilosity. Two anepisternal bristles. Scutellum yellow with apical markings black; basally with two distinct dark spots. Legs dark yellow to orange brown; setation typical of subgenus. Setae dark, front femur with posterior row dark. Wings with brownish bands, yellow markings strongly reduced. Banding, setation and venation normal for subgenus. Marginal band continuous; discal band joined with marginal band; cross-vein r-m at middle of discal cell; vein R1 ending before or oppostie cross-vein r-m. Abdomen. greyish, with clearly defined spots. Pattern of spots and setation normal for subgenus.
Female
As male except for the following characters. Third antennal segment two to three times as long as second. Frons yellow, not silvery. Face wholly yellow; at antennal socket sometimes with a darker horizontal band. Oviscape shorter than abdominal terga 3-6, orange in colour.
(Description after De Meyer, 1996)
See description of Ceratitis punctata (Wiedemann, 1824) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis quinaria (Bezzi, 1918)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) quinaria (Bezzi)
Pardalaspis quinaria Bezzi, 1918: 235.
Body length: 4.20 (3.60-4.75) mm; wing length: 3.93 (3.50-4.50) mm.

Male
Head. Antenna yellow to yellow-orange. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons pale yellow, centre usually deeper yellow coloured; with short scattered hairs which are largely the same colour as frons; concave to flat, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus, except more reddish. Thorax. Postpronotum white or white-yellow, unspotted. Ground colour of mesonotum pale; mesonotal pattern variable, dorsocentral spots often less distinct. Chaetotaxy normal for subgenus. Scapular setae pale. One anepisternal bristle. Scutellum white basally, otherwise yellow with three apical and two subapical small black spots; basally without dark spots, only slight darker colouration. Subscutellum completely pale. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing bands with colour of markings variable, yellow to yellow-brown. Banding, marginal band continuous; cubital band free; medial band absent; crossvein r-m well before middle of discal cell, usually at basal third. Crossvein dm-cu position variable. Abdomen. Pale yellow colour. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Scutellum in Aden specimen with lateral sets of spots partly fused. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis quinaria (Bezzi, 1918) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis rosa Karsch, 1887"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) rosa Karsch
Ceratitis rosa Karsch, 1887: 1.
Body length: 4.96 (4.25-5.30) mm; wing length: 5.34 (4.50-5.75) mm.

Male
Head:
Antenna yellow. First flagellomere 2-3 times as long as pedicel. Arista with short to moderately long rays; ventral rays shorter and sparser than dorsal rays, especially basally. Frons yellow; with short scattered setulae distinctly darker than frons. Frontal setae well developed. Face yellowish white. Genal seta and setulae dark, well developed. Thorax:
Postpronotal lobe yellowish white, without spot, although sometimes darker yellow around postpronotal seta. Scutal pattern:
ground color grayish- brown with orange tinge; with streaks and darker markings but without distinct spots except prescutellar white markings separate, usually with paler area in between. Scapular setae dark. Scutellum yellowish white, basally usually with two separate dark spots, sometimes less distinct; apically with three separate black spots, extending to basal 0.33. Anepisternum on ventral half darker yellowish brown; setulae pale. Legs:
Yellow except where otherwise noted; setation typical for subgenus, mainly pale. Foreleg:
femur without bushy feathering posteriorly, only dispersed rows of long black setulae posterodorsally, posteroventrally shorter and pale; ventral setae black. Midleg:
femur with few dispersed pale setulae ventrally; tibia moderately broadened; anteriorly black with conspicuous silvery shine when viewed from certain angle on distal 0.66 to 0.75 (black color sometimes inconspicuous in teneral specimens but silvery shine is always present) with black feathering dorsally along distal 0.75 and ventrally along distal 0.66, ocassionaly to distal 0.75. Hindleg:
femur at apical 0.25 with longer setulae dorsally and ventrally. Wing:
banding yellowish brown. Interruption between marginal and discal bands near vein R1 clear and complete; cubital band free; medial band absent; crossvein R-M opposite middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu oblique anterobasally. Abdomen:
Mostly yellow. Tergites 2 and 4 with pale gray band on posterior half, anterior margin sometimes with narrowly brownish colored, especially laterally. Tergite 3 with posterior half patchily brownish colored, anterior half yellowish brown, both parts not clearly demarcated; sometimes more complete brown. Tergite 5 with basal half brownish, sometimes divided medially into two spots.
Male epandrium in lateral view with posterior lobe of lateral surstylus short and straight, anterior lobe well pronounced.
Female
As male except for the following characters:
First flagellomere yellowish orange. Crossvein DM-Cu oblique posterobasally. Anepisternum on ventral part rarely with darker setulae. Legs without feathering; forefemur posteroventrally with pale pilosity, at least basally. Oviscape shorter than preabdomen. Aculeus at most six times longer than wide; tip with distinct apical indentation and lateral margin slightly sinuous (specimens from Luabo, Mozambique seem to have an aculeus tip which is without any indentation but otherwise completely fit the description of rosa. They are tentatively placed here).
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis rosa Karsch, 1887 in source PDF."""
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Marc de Meyer

"""Ceratitis roubaudi (Bezzi, 1923)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pterandrus) roubaudi (Bezzi)
Pardalaspis roubaudi Bezzi, 1923: 527.
Body length: 5.13 (4.40-6.15) mm; wing length: 5.04 (4.60-5.40) mm.

Male
Head:
Antenna orange to brown (yellow according to the original description). First flagellomere twice as long as pedicel. Arista with short rays (medium long rays according to the original description); ventral rays shorter and sparser than dorsal rays, especially basally. Frons convex; dark yellow, with short scattered setulae distinctly darker than frons. Face white; gena darker yellow. Genal seta and setulae dark, well developed. Thorax:
Postpronotal lobe white (yellow according to the original description), with brown spot. Scutal pattern, ground color dark brown, microtrichose areas silvery with ashgray shine; prescutellar white markings separate, with pale gray area in between. Scapular setae dark. Scutellum white, basally with two separate dark spots (not mentioned in original description), apically with three separate black spots, reaching to basal 0.3 of scutellum. Anepisternum yellow, ventral half brown; one anepisternal seta; anepisternal setulae pale, except in ventral half. Legs:
Yellowish brownish (yellow according to original description); setation typical for subgenus, mainly black; no feathering. Wing:
Bands brown. Marginal band forming continuous band with anterior part of discal band; cubital band free; medial band absent; crossvein R-M just proximal of middle of discal cell. Apex of vein R1 distal to level of crossvein R-M. Crossvein DM-Cu oblique anterobasally. Abdomen:
Mostly pale orange. Tergites 2 and 4 with gray transverse band along posterior half, brownish anteriorly. Tergite 3 largely brown. Setation typical for subgenus.
Male terminalia not dissected.
Female
As male except for the following characters:
One or two anepisternal setae present. Crossvein R-M opposite middle of discal cell. Oviscape shorter than preabdomen. Aculeus about six times longer than wide; tip pointed and lateral margin slightly concave.
(Description after De Meyer & Freidberg, 2006)
See description of Ceratitis roubaudi (Bezzi, 1923) in source PDF."""
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Marc de Meyer

"""Ceratitis scaevolae (Munro, 1929)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) scaevolae (Munro)
Pardalaspis scaevolae Munro, 1929: 394.
Body length: 5.14 (4.60-6.25) mm; wing length: 4.36 (3.85-4.75) mm.

Male
Head. Antenna yellow to yellow-orange. Third antennal segment twice as long as second segment. Arista bare. Frons with short scattered hairs which are largely the same colour as the frons; convex, in lateral view slightly projecting forwards at antennal implant. Face usually yellow-white. Chaetotaxy normal for subgenus. Thorax. Postpronotum white, unspotted. Ground colour of mesonotum pale with slight orange tinge. Chaetotaxy normal for subgenus. Scapular setae usually black, sometimes yellow-reddish (inner) to reddish black. One anepisternal bristle. Scutellum white, with three black markings joined apically; basally no spots. Subscutellum completely dark. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur darkish. Wing bands with markings extensively yellow to yellow-brown. Banding, marginal band continuous; cubital band joined with discal band; medial band absent; crossvein r-m well beyond middle of discal cell, vein R1 before crossvein. Crossvein dm-cu posterioly more outwards than anteriorly. Abdomen. Yellow colour. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Posterior and posterodorsal rows on fore femur sometimes less dark; scutellar spots occasionally joined together, forming one large semicircular spot. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis scaevolae (Munro, 1929) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis semipunctata De Meyer, 1996"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pardalaspis) semipunctata De Meyer
Ceratitis (Pardalaspis) semipunctata De Meyer, 1996 : 24.
Body length: 7.25 (7.0-7.5) mm; wing length: 7.5 mm.

Female
Head. Antennal segments orange. Third antennal segment twice to three times as long as second segment. Arista basal part pale, otherwise dark. Frons pale yellowish in ground color, silvery over entire length; with dispersed short hairs, distinctly darker than frons. Ocellar triangle dark. Face pale yellow. Occiput moderately swollen below, pale whitish. Chaetotaxy normal for subgenus. Thorax. Ground colour of mesonotum pale; longitudinal lines hardly discernible. Postpronotum same colour as mesonotum. Chaetotaxy normal for subgenus. Mesonotum with pale pilosity. Anepisternum along upper margin with darker hair, and a few dark hairs on lower part of anepisternum. Two or three anepisternal bristles. Scutellum yellow with apical markings black; basally without dark spots, only a faint darker coloration. Legs pale yellow; setation typical of subgenus. Front femur predominantly with pale setae, posterior row dark. Wings with brownish bands, yellow markings strongly reduced. Banding, setation and venation normal for subgenus. Marginal band continuous; discal band joined with marginal band; cross-vein r-m at middle of discal cell; vein R1 ending before cross-vein r-m. Abdomen. Greyish, with clearly defined spots. Pattern of spots and setation normal for subgenus. Oviscape shorter than abdominal terga 3-6, yellow-orange in colour.
Male
Unknown.
(Description after De Meyer, 1996)
See description of Ceratitis semipunctata De Meyer, 1996 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis serrata De Meyer, 1996"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pardalaspis) serrata De Meyer
Ceratitis (Pardalaspis) serrata De Meyer, 1996 : 24.
Body length: 6.5 mm; wing length: 6.0 mm.

Female
Head. Antennal segments missing. Frons yellow in ground color, weakly silvery in the middle; with dispersed short hairs, darker than frons. Ocellar triangle dark. Face yellow-white, no median band. Occiput moderately swollen below, pale whitish. Chaetotaxy normal for subgenus. Thorax. Ground colour of mesonotum with an orange tinge; with darker coloration around setal sockets and some darker streaking, not clearly defined. Postpronotum same colour as mesonotum. Chaetotaxy normal for subgenus. Mesonotum with pale pilosity. Anepisternum with darker hairs over entire surface. Three anepisternal bristles. Scutellum yellow with apical markings black; basally with two dark spots. Legs yellow; setation typical of subgenus. Hind femur with dorsal hairs dark apically. Setae dark, posterior row on front femur dark. Wings with brownish bands, yellow markings strongly reduced. Banding, setation and venation normal for subgenus. Marginal band continuous; discal band joined with marginal band; cross-vein r-m at middle of discal cell; vein R1 ending before cross-vein r-m. Abdomen. Grey-brown, with orange tinge and clearly defined spots. Pattern of spots and setation normal for subgenus. Oviscape short, shorter than abdominal terga 3-6.
Male
Unknown.
(Description after De Meyer, 1996)
See description of Ceratitis serrata De Meyer, 1996 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis silvestrii Bezzi, 1912"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) silvestrii Bezzi
Ceratitis silvestrii Bezzi, 1912: 10.
Body length: 3.88 (3.75-4.00) mm; wing length: 3.96 (3.75-4.15) mm.

Male
Head. Antenna yellow. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons with short scattered hairs which are the same colour as frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Face white. Chaetotaxy normal for subgenus, except dark reddish in colour. Thorax. Postpronotum white, unspotted. Ground colour of mesonotum almost white. Chaetotaxy normal for subgenus, dark reddish. Scapular setae pale. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically; basally without spots. Subscutellum completely dark. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Ventral bristles reddish. Wing bands with yellow markings. Banding, marginal band continuous; cubital band free, medial band absent; crossvein r-m well before middle of discal cell, at basal third. Crossvein dm-cu posteriorly approximately at same position as anteriorly. Abdomen. Yellow-grey. Setation and banding typical for subgenus.
Female
As in male except for the following characters:
Mesonotum less white.
(Description after De Meyer, 1998)
See description of Ceratitis silvestrii Bezzi, 1912 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis simi Munro, 1933"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) simi Munro
Ceratitis (Pardalaspis) simi Munro, 1933a: 37.
Body length: 6.38 (5.40-7.15) mm; wing length: 6.24 (5.85-7.25) mm.

Male
Head. Antenna yellow to yellow-orange. Third antennal segment twice as long as second segment. Arista with very short hairs over entire length. Frons with short scattered hairs which are distinctly darker than the frons; concave to flat, in lateral view slightly projecting forwards at antennal implant. Face usually yellow-white, but sometimes somewhat darker, more yellow. Darker patches at lower eye margin, but not always outspoken. Chaetotaxy normal for subgenus. Thorax. Postpronotum white or white-yellow, spotted. Ground colour of mesonotum pale with orange tinge. Chaetotaxy normal for subgenus. Scapular setae black. One or two anepisternal bristles. Scutellum white basally, otherwise yellow with three black separate markings apically; basally two dark but inconspicuous spots. Subscutellum pale with three dark spots, touching along dorsal margin. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing bands with markings extensively yellow. Banding, marginal band continuous; cubital band joined with discal band; medial band free; crossvein r-m at middle of discal cell. Crossvein dm-cu position variable. Abdomen. Dark yellow colour. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Scutellum with basal spots more outspoken. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis simi Munro, 1933 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis stictica Bezzi, 1909"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) stictica Bezzi
Ceratitis stictica Bezzi, 1909: 277.
Body length: 4.93 (4.70-5.30) mm; wing length: 4.93 (4.50-5.20) mm.

Male
Head. Antenna yellow-orange. Third antennal segment three times as long as second segment. Arista with medium long hairs over entire length. Frons yellow-orange, with short scattered hairs which are largely the same colour as frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Frons and face sometimes with brownish median spots, no distinct bands. Thorax. Postpronotum yellow-white, spotted. Ground colour of mesonotum pale with yellow-orange to orange tinge. Chaetotaxy normal for subgenus. Scapular setae pale (occasionally more darkish). One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically; basally with two dark separate spots. Subscutellum with three separate spots, sometimes joined dorsally. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Ventral spines on fore femur with pale bristles, occasionally dark. Wing bands with extensively yellow markings. Banding variable, marginal band continuous; cubital band usually weakly joined with discal band, sometimes free; medial band indistinct and/or free, sometimes joined with marginal band; crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus.
Female. As male except for the following characters:
Subscutellum sometimes completely black. Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis stictica Bezzi, 1909 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis striatella Munro, 1935"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) striatella Munro
Ceratitis (Pardalaspis) striatella Munro, 1935: 310.
Body length: 6.64 (5.00-7.50) mm; wing length: 6.21 (5.80-6.90) mm.

Male
Head. Antenna yellow-orange. Third antennal segment three times as long as second segment. Arista with short hairs over entire length. Frons yellow to orange; with short scattered hairs which are distinctly darker than the frons; concave to flat, in lateral view not distinctly projecting forwards at antennal implant. Face yellow-orange. Darker, more orange patches at lower eye margin, continued along lower margin of face. Chaetotaxy normal for subgenus. Thorax. Postpronotum yellow, slightly paler than ground colour of mesonotum, with black spot. Ground colour of mesonotum mainly yellow-orange, sometimes more greyish. Chaetotaxy normal for subgenus. Scapular setae pale (in holotype of striatella black except one inner seta pale). One or two anepisternal bristles. Scutellum yellow with three black separate markings apically; basally two dark separate spots. Subscutellum pale with three dark spots, widely separate. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Ventral spines black. Wing bands with markings extensively yellow. Banding, marginal band continuous; cubital band free (at most weakly connected with discal band); medial band absent; crossvein r-m before middle of discal cell. Crossvein dm-cu posteriorly more outwards than anteriorly. Abdomen. Yellow colour. Setation and banding typical for subgenus.
Female. As male except for the following characters:
Frons sometimes weakly projected forwards at antennal implant. Crossvein r-m variable. Oviscape very long, at least twice as long as abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis striatella Munro, 1935 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis sucini De Meyer, 1998"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) sucini De Meyer
Ceratitis (Ceratalaspis) sucini De Meyer, 1998 : 287.
Body length: 5.85 mm; wing length: 5.40 mm.

Female
Head. Antenna yellow-orange. Third antennal segment two times as long as second segment. Arista with short hairs over entire length. Frons yellow; with short scattered hairs which are largely the same colour as the frons; in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum white; unspotted. Ground colour of mesonotum pale. Chaetotaxy normal for subgenus. Scapular setae pale. One anepisternal bristle. Scutellum yellow-white; with three black separate markings; basally without distinct spots. Subscutellum pale with three dark separate spots. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Ventral spines on fore femur reddish. Wing bands with markings extensively yellow. Banding, marginal band continuous; cubital band free; medial band joined with marginal band; crossvein r-m just before middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Dark yellow. Setation and banding typical for subgenus. Oviscape shorter than abdomen.
Male
Unknown.
(Description after De Meyer, 1998)
See description of Ceratitis sucini De Meyer, 1998 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis turneri (Munro, 1937)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) turneri (Munro)
Pardalaspis turneri Munro, 1937: 5.
Body length: 3.85 (3.75-4.25) mm; wing length: 3.49 (3.85-4.35) mm.

Male
Head. Antenna yellow-orange except third segment brownish. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Face with upper half darker coloured. Frons with short scattered hairs which are largely the same colour as frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum white, unspotted. Ground colour of mesonotum pale, partly with orange tinge (not distinct because of broad dark bands); mesonotal pattern, spots usually confluent forming two broad lateral stripes with white lunule, sometimes less developed. Chaetotaxy normal for subgenus. Scapular setae black, sometimes inner setae pale. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically, not extending more than halfway scutellum (some specimens from Kenya have the spots partly joined); basally without dark spots, only slight darker colouration. Subscutellum completely dark. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing bands with yellow areas reduced, mainly dark. Banding, marginal band with clear interruption after cell R1; cubital and medial bands joined with marginal band; crossvein r-m before middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Extension cup sometimes weakly sinoid. Abdomen. Brownish colour. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Oviscape shorter than abdominal terga 3-6 combined.
(Description after De Meyer, 1998)
See description of Ceratitis turneri (Munro, 1937) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis venusta (Munro, 1956)"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Ceratalaspis) venusta (Munro)
Hoplolophomyia venusta Munro, 1956: 466.
Body length: 5.60 (5.10-5.85) mm; wing length: 5.77 (5.40-5.90) mm.

Male
Head. Antenna yellow. Third antennal segment twice as long as second segment. Arista with short hairs over entire length. Frons with short scattered hairs which are largely the same colour as frons; convex, in lateral view not distinctly projecting forwards at antennal implant. Chaetotaxy normal for subgenus. Thorax. Postpronotum white, unspotted. Ground colour of mesonotum pale with orange tinge; mesonotal pattern, median spot between postpronota usually absent, sometimes present as darker patch. Chaetotaxy normal for subgenus. Scapular setae black. One anepisternal bristle. Scutellum white basally, otherwise yellow with three black separate markings apically, hardly visible in dorsal view, restricted to apical margin; basally without dark spots, only slight darker colouration. Subscutellum completely dark. Legs yellow; setation typical for subgenus, mainly pale especially on femora. Posterior and posterodorsal rows on fore femur pale. Wing bands markings extensively yellow. Banding, marginal band with clear interruption after cell R1; cubital and medial bands joined with marginal band (exceptionaly the cubital band is separate); crossvein r-m at middle of discal cell. Crossvein dm-cu posteriorly more inwards than anteriorly. Abdomen. Yellow. Setation and banding typical for subgenus.
Female
As male except for the following characters:
Ocellar bristles sometimes shorter, at most 1.5 times length of ocellar triangle. Occasionally only one frontal bristle. Lower eye margin sometimes with darker colouration. Posterior and posterodorsal rows on fore femur apically with dark hairs. Crossvein r-m before middle of discal cell. Oviscape shorter than abdominal terga 3-6 combined (in one specimen longer than terga). Aculeus with tip variable.
(Description after De Meyer, 1998)
See description of Ceratitis venusta (Munro, 1956) in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

"""Ceratitis zairensis De Meyer, 1996"""

provided by Afrotropical Fruitfly Project
"""Ceratitis (Pardalaspis) zairensis De Meyer
Ceratitis (Pardalaspis) zairensis De Meyer, 1996 : 24.
Body length: 7.0 mm; wing length: 7.0 mm.

Male
Head. Antennal segments orange. Third antennal segment twice as long as second segment. Arista with basal part orange coloured, otherwise dark. Frons dark yellow, silvery over entire length but not distinct especially not in centre; with scattered short hairs, darker than ground colour. Ocellar triangle dark. Face yellow-orange; no median band. Occiput moderately swollen below, pale whitish. Chaetotaxy normal for subgenus. Thorax. Ground colour of mesonotum pale with orange tinge; no distinct spots, except around sockets of dorsocentral and prescutellar acrostichal setae, these markings indistinct. Postpronotum same colour as centre of mesonotum. Chaetotaxy normal for subgenus. Mesonotum with pale pilosity. Anepisternum with pale pilosity except upper margin with darker hairs. Three anepisternal bristles, median one less developed than other two. Scutellum white-yellow with apical markings black; basally without distinct dark spots, but clearly darker coloration. Legs yellow; setation typical of subgenus; predominantly pale, posterior row on fore femur dark. Wings with bands mainly yellowish-brown. Banding, setation and venation normal for subgenus. Marginal band continuous; discal band joined with marginal band; cross-vein r-m before middle of discal cell; vein R1 ending beyond cross-vein r-m. Abdomen. Grey-brown with orange tinge, with weakly defined spots. Pattern of spots and setation normal for subgenus.
Female
Unknown.
(Description after De Meyer, 1996)
See description of Ceratitis zairensis De Meyer, 1996 in source PDF."""
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"""Marc de Meyer, MRAC"""
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Marc de Meyer

Dier ( Afrikaans )

provided by wikipedia AF

Diere behoort tot Animalia, een van vyf (of soms ses) biologiese koninkryke. Dit word ook Metazoa genoem. Diere is oor die algemeen veelsellige, eukariotiese organismes wat ander organismes eet vir voeding. Die meeste diere kan in die een of ander stadium van hul lewe onafhanklik en spontaan beweeg.

Diere bestaan reeds van die Prekambrium (PreЄ) af, maar die meeste bekende dierefilums het tydens die Kambriese ontploffing, sowat 542 miljoen jaar gelede, in die fossielrekord verskyn. Diere kan oor die algemeen in gewerweldes en ongewerweldes verdeel word. Gewerweldes het ’n ruggraat en maak minder as 5% van alle dierespesies uit. Dit sluit in visse, amfibieë, reptiele, voëls en soogdiere. Die res is ongewerweldes; hulle het nie ’n ruggraat nie. Dit sluit in weekdiere (mossels, oesters, seekatte, pylinkvisse, slakke); geleedpotiges (insekte, spinnekoppe, skerpioene, krappe, krewe, garnale); ringwurms (erdwurms, bloedsuiers), rondewurms, platwurms, neteldiere (jellievisse, seeanemone, korale), ribkwalle en sponsdiere.

Die studie van diere word dierkunde of soölogie genoem.

Etimologie

Die naam "Animalia" kom van die Latynse animalis, wat beteken "het asem", "het ’n siel" of "lewende wese".[1] Die Afrikaanse woord "dier" sluit oor die algemeen mense uit – gewoonlik word soogdiere en ander gewerweldes bedoel. Die biologiese definisie verwys na alle lede van die koninkryk Animalia – so uiteenlopend soos sponse, jellievisse, insekte en mense.[2]

Geskiedenis van klassifikasie

 src=
Carl Linnaeus is bekend as die vader van moderne taksonomie.[3]

Aristoteles het die lewende wêreld in diere en plante verdeel, en Carl Linnaeus het die eerste hiëragiese klassifikasie ingestel.[4] In Linnaeus se oorspronklike skema was diere een van drie koninkryke, en dit is verdeel in Vermes, Insecta, Pisces, Amphibia, Aves en Mammalia. Sedertdien is die laaste vier verenig onder die filum Chordata.

In 1874 het Ernst Haeckel die diereryk in twee subkoninkryke verdeel: Metazoa (meerselliges) en Protozoa (eenselliges).[5] Die Protozoa is later na die koninkryk Protista geskuif, en Metazoa word nou beskou as ’n sinoniem vir Animalia.[6]

Eienskappe

Diere het verskeie eienskappe wat hulle van ander lewensvorme onderskei. Hulle is eukarioties en meersellig,[7] wat hulle onderskei van bakterieë en die meeste Protista-organismes. Hulle is heterotrofies (verorber ander organismes vir voeding)[8] en verteer kos gewoonlik in ’n interne kamer, wat hulle onderskei van plante en alge.[9] Hulle verskil ook van plante, alge en swamme deurdat hulle nie rigiede selwande het nie.[10]

Alle diere is beweeglik,[11]al is dit net in sekere lewenstadiums. By die meeste gaan embrio's deur ’n blastulastadium[12] (’n kiemblasiestadium wanneer die embrio die vorm het van 'n sfeer waarvan die wand uit een sellaag bestaan),[13] ’n eienskap wat eie aan diere is.

Struktuur

Met ’n paar uitsonderings, soos die sponse, het diere liggame wat uit verskillende soorte weefsel bestaan. Dit sluit in spiere, wat kan saamtrek en beweging beheer, en senuwees, wat seine stuur en verwerk. Daar is ook gewoonlik ’n interne spysverteringskamer met een of twee openinge.[14]

Alle diere het eukariotiese selle, omring deur ’n kenmerkende ekstrasellulêre matriks, wat bestaan uit kollageen en elastiese glukoproteïene.[15] Dit kan verkalk wees om strukture soos skulpe, bene en sponsnaalde te vorm.[16] Tydens ontwikkeling vorm dit ’n relatief buigbare raamwerk[17] waarop selle kan rondbeweeg en reorganiseer kan word, en dit maak ingewikkelde strukture moontlik. In teenstelling hiermee het ander meersellige organismes, soos plante en swamme, selle wat op plek gehou word deur selwande, en hulle ontwikkel so deur progressiewe groei.[14]

Voortplanting en ontwikkeling

 src=
Sommige spesies landslakke gebruik "liefdespyltjies" as ’n vorm van seksuele seleksie.[18]

Byna alle diere is gewikkel in die een of ander vorm van seksuele voortplanting.[19] Hulle vervaardig haploïede gamete deur meiose. Die kleiner, beweeglike gamete is spermselle en die groter, onbeweeglike gamete is eierselle.[20] Hulle smelt saam en vorm sigote, wat in nuwe individue ontwikkel.[21]

Baie diere kan ook aseksueel voortplant.[22] Dit kan geskied deur middel van partenogenese, waar bevrugte eierselle geproduseer word sonder paring, knopvorming of verdeling.[23]

 src=
Manlike sjimpansees bly dikwels in die gemeenskap waarin hulle gebore word, terwyl wyfies by ander groepe aansluit.

’n Sigoot ontwikkel aanvanklik in ’n hol sfeer, wat ’n blastula genoem word;[24] dit ondergaan herrangskikking en differensiasie. In sponse swem blastulalarwes na ’n nuwe ligging en ontwikkel in ’n nuwe spons.[25] In die meeste ander groepe ondergaan die blastula ’n meer ingewikkelde herrangskikking.[26] Dit vorm eers ’n invaginasie en dan ’n gastrula met ’n spysverteringskamer en twee aparte kiemlae – ’n eksterne ektoderm en ’n interne endoderm.[27] In die meeste gevalle ontwikkel daar ook ’n mesoderm tussenin.[28] Hierdie kiemlae differensieer dan om weefsel en organe te vorm.[29]

Vermyding van inteling

By seksuele voortplanting lei inteling (met ’n nabye familielid) gewoonlik tot ’n afname in biologiese gehalte, soos die voorkoms van ’n vroeë dood by sekere spesies.[30] Diere het verskeie maniere ontwikkel om inteling te vermy en uitkruising te bevorder.[31] Sjimpansees gebruik byvoorbeeld verspreiding om te voorkom dat nabye familielede paar;[31] individuele diere versprei na verskillende groepe en weg van die groep waarin hulle gebore is.

In verskeie spesies paar wyfies met verskillende mannetjies en bring dus kleintjies voort van ’n hoër genetiese gehalte. Dit gebeur veral by wyfies wat ’n paartjie gevorm het met ’n mannetjie van swak genetiese gehalte, soos ’n nabye familielid.[32]

Kos- en energiebronne

Diere is heterotrofies, wat beteken hulle eet ander organismes, direk of indirek.[33] Hulle word gewoonlik ook verder verdeel in groepe soos karnivore, herbivore, omnivore en parasiete.[34]

Predasie is ’n biologiese interaksie waar ’n roofdier (’n heteretroof wat jag) sy prooi (die organisme wat aangeval word) eet.[35] ’n Roofdier het dalk of dalk nie sy prooi doodgemaak voordat hy dit eet, maar die daad van predasie lei gewoonlik daartoe dat die prooi doodgemaak word.[36] Die ander belangrike kosbron is dooie organiese materiaal.[37] Dit is soms moeilik om tussen dié twee voedingsvorme te onderskei – soos wanneer ’n parasiet ’n gas-organisme eet en dan sy eiers daarop lê sodat sy kleintjies die verrottende karkas kan eet. Druk wat hulle op mekaar uitoefen, lei dikwels tot ’n evolusie-resies tussen die jagter en prooi, en dit lei tot verskeie evolusie-aanpassings in albei.[38]

Die meeste diere gebruik die energie van sonlig direk of indirek deur plante of plantvretende diere te eet. Fotosintese in plante omskep die energie van sonlig in chemiese energie in die vorm van eenvoudige suikers (byvoorbeeld glukose). Hierdie suikers word dan gebruik as die boustene vir plantgroei.[14] Wanneer ’n dier die plante vreet, of diere vreet wat die plante gevreet het, verkry hulle hul energie daaruit. Die energie help die dier groei en gee hom die energie om te beweeg.[39][40]

Diere wat naby hidrotermiese of koue bronne op die seebodem voorkom, is nie van die energie van sonlig afhanklik nie.[41] In plaas daarvan vorm chemosintetiese Archaea en bakterieë die basis van die voedselketting.[42]

Oorsprong en fossielrekord

 src=
Dunkleosteus was ’n 10 m lange prehistoriese vis.[43]

Daar word algemeen geglo diere het binne die groep eukariote met sweephare (flagella) ontstaan.[44] Hul naaste bekende verwante wat steeds bestaan, is die Choanoflagellatea (kraagsweepdiertjies) waarvan die morfologie ooreenstem met dié van die koanosiete (kraagselle) van sekere sponse.[45]

Molekulêre studies plaas diere in ’n supergroep genaamd Opisthokonta, wat ook Choanoflagellatea, die swamme en ’n paar klein parasitiese protiste insluit.[46] Die naam kom van die ligging van die flagellum aan die agterkant van beweeglike selle, soos by die meeste diere se spermselle, terwyl ander eukariote se flagellum gewoonlik aan die voorkant is.[47]

Die eerste fossiele wat dalk dié van diere is, is uit die Proterosoïese Eon en het voorgekom in 665 miljoen jaar oue rotse[48] in die Trezona-formasie van Suid-Australië.[48] Daar word geglo die fossiele was van vroeë sponsdiere.

Die volgende moontlike dierefossiele was uit die einde van die Prekambrium, sowat 610 miljoen jaar gelede, en is bekend as die Ediacariese biota.[49] Dit is egter moeilik om ’n verband te vind met latere fossiele. Sommige was dalk voorlopers van moderne filums, hulle kon aparte groepe gewees het of was dalk glad nie diere nie.[50]

Die meeste bekende filums maak hul verskyning min of meer op dieselfde tyd in die Kambrium-periode, sowat 542 miljoen jaar gelede.[51] Dit is steeds onbekend of dié gebeurtenis, bekend as die Kambriese ontploffing, veroorsaak is deur ’n vinnige differensiasie tussen verskillende groepe en of dit was vanweë ’n verandering in toestande wat fossilering moontlik gemaak het.

Sommige paleontoloë dink diere het lank voor die Kambriese ontploffing ontstaan, dalk reeds ’n miljard jaar gelede.[52] Spoorfossiele soos voetspore en gate uit die Tonium-periode wat ontdek is, dui op die moontlike teenwoordigheid van tripoblastiese wurms (met drie kiemlae), omtrent so groot (sowat 5 mm breed) en kompleks soos erdwurms.[53]

Aan die begin van die Tonium sowat ’n miljard jaar gelede was daar ’n afname in die verskeidenheid van stromatoliete, wat dalk daarop dui dat weidiere hul verskyning gemaak het aangesien stromatoliet-verskeidenheid toegeneem het nadat weidiere uitgesterf het met die Perm-Trias- en die Ordovisium-Siluur-uitwissing. Dit het weer afgeneem kort nadat die weidierbevolking herstel het. Die ontdekking dat soortgelyke spore vandag geskep word deur die reusagtige, eensellige protis Gromia sphaerica, werp egter ’n skadu oor die vertolking van die fossiele as bewyse van ’n vroeë evolusie van diere.[54][55]

Groepe diere

Beide tradisionele morfologiese en moderne molekulêre filogenetiese ontledings dui op ’n groot evolusionêre oorgang van diere sonder ’n bilaterale simmetrie (Porifera, Ctenophora, Cnidaria en Placozoa) na dié wat wel bilateraal simmetries is (Bilateria). Laasgenoemde word verder geklassifiseer in óf Deuterostomia (waarby die eerste opening van die spysverteringstelsel die anus word en die tweede een die mond) óf Protostomia (waarby die mond eerste gevorm word). Die verwantskap tussen diere sonder ’n bilaterale simmetrie word betwis, maar alle diere met ’n bilaterale simmetrie vorm vermoedlik ’n monofiletiese groep. Die huidige begrip van die verwantskap tussen die groot groepe diere word deur die volgende kladogram opgesom:[56]


Apoikozoa

ChoanoflagellateaCronoflagelado2.svg


Animalia

Porifera



Placozoa



CtenophoraMertensia ovum.png



CnidariaMedusae of world-vol03 fig360 Atolla chuni.jpg



Bilateria

DeuterostomiaCyprinus carpio3.jpg


Protostomia

EcdysozoaAcrodipsas brisbanensis.jpg



LophotrochozoaLoligo forbesii.jpg









Diere sonder bilaterale simmetrie

Verskeie dierefilums word gekenmerk deur die afwesigheid van bilaterale simmetrie en daar word geglo hulle het vroeg in die evolusie uit ander diere ontwikkel. Die sponse (Porifera) het vermoedelik eerste ontwikkel en word beskou as die oudste dierefilum.[57] Hulle het nie die komplekse organisasie wat in die meeste ander filums aangetref word nie.[58] Hul selle is gedifferensieer, maar in die meeste gevalle nie in aparte soorte weefsel georganiseer nie.[59] Sponse neem gewoonlik voedingstowwe in deur water deur gaatjies in te trek.[60] ’n Reeks studies van 2008 tot 2015 ondersteun egter die mening dat Ctenophora, of die ribkwalle, die basale lyn van diere is.[61][62][63] Dié bevinding is omstrede, want dit dui daarop dat sponse dalk nie so primitief is nie,[61] en ander navorsers meen dit is ’n statistieke ongerymdheid.[64][65][66]

 src=
’n Oranje olifantoorspons, Agelas clathrodes, voor en twee korale, Iciligorgia schrammi en Plexaurella nutans, agter.

Filums soos Ctenophora en Cnidaria, wat seeanemone, korale en jellievisse insluit, is radiaal (straalsgewys) simmetries en het spysverteringskamers met een opening wat as mond én anus dien.[67] Albei het verskillende soorte weefsel, maar dit is nie in organe georganiseer nie.[68] Daar is net twee hoofkiemlae, die ektoderm en endoderm, met net verstrooide selle tussenin. Daarom word dié diere soms "diploblasties" genoem.[69]

Daar word nou geglo die Myxozoa, mikroskopiese parasiete wat aanvanklik as Protozoa beskou is, het binne Cnidaria ontwikkel.[70]

Diere met bilaterale simmetrie

Alle ander diere vorm ’n monofiletiese groep wat Bilateria genoem word. Hulle is meestal bilateraal simmetries en het dikwels ’n gespesialseerde kop met voedings- en sintuiglike organe. Die liggaam is triploblasties – dit beteken al drie kiemlae is goed ontwikkel – en weefsel vorm afsonderlike organe. Die spysverteringskanaal het twee openinge, ’n mond en ’n anus, en daar is ’n interne holte bekend as ’n seloom (of buikholte) of pseudoseloom (skynbuikholte). Daar is uitsonderings op al hierdie eienskappe.

Genetiese studies het ons begrip van die verhouding tussen die diere in Bilateria aansienlik verander. Dit lyk of die meeste tot twee groot stambome behoort: Deuterostomia en Protostomia; laasgenoemde sluit in die Ecdysozoa en Lophotrochozoa. Die Chaetognatha is aanvanklik as Deuterostomia geklassifiseer, maar onlangse molekulêre studies het dié groep geëien as ’n basale Protostomia-lyn.[71]

Daar is ook ’n paar klein groepe diere met bilaterale simmetrie met relatief kriptiese morfologie waarvan die verwantskap met ander diere nog nie baie duidelik is nie. So het onlangse molekulêre studies Acoelomorpha en Xenoturbella geëien as behorende tot ’n monofiletiese groep,[72][73][74] maar daar is weinig ooreenstemming oor of die groep binne Deuterostomia ontwikkel het,[73] en of dit ’n sustergroep van alle ander diere met bilaterale simmetrie (Nephrozoa) verteenwoordig.[75][76] Ander groepe met onsekere verwantskap sluit Rhombozoa en Orthonectida in. Een filum, Monoblastozoa, is in 1892 deur ’n wetenskaplike beskryf, maar tot dusver was daar geen verdere bewys dat dit bestaan nie.[77]

Deuterostomia en Protostomia

 src=
Twee winterkoninkies van die spesie Malurus cyaneus.

Deuterostomia verskil in verskeie opsigte van Protostomia. Diere van albei groepe het ’n volledige spysverteringstelsel. By laasgenoemde groep ontwikkel die eerste opening wat in die embrio verskyn, in die mond en die tweede opening in die anus. In eersgenoemde groep ontwikkel die openinge andersom.[78] By die meeste Protostomia vul selle eenvoudig die binnekant van die gastrula in om die mesoderm te vorm, maar by Deuterostomia vorm dit deur die invaginasie van die endoderm.[79] Deuterostomia-embrio's ondergaan radiale kliewing (verdeling) en Protostomia spirale kliewing.[80]

Dit alles dui daarop dat Deuterostomia en Protostomia twee aparte, monofiletiese stambome is. Die belangrikste filums van Deuterostomia is Echinodermata en Chordata.[81] Eersgenoemde is radiaal simmetries en uitsluitlik seediere, soos seesterre, seekastaiings en seekomkommers.[82] Chordata word deur die gewerweldes, diere met rugstringe, oorheers.[83] Dit sluit in visse, amfibieë, reptiele, voëls en soogdiere.[84]

Deuterostomia sluit ook Hemichordata, of die kraagdraers, in.[85][86] Hoewel hulle nie vandag juis volop is nie, behoort die belangrike fossiele graptoliete dalk tot hierdie groep.[87]

Ecdysozoa

 src=
Geelvlerknaaldekokers, Sympetrum flaveolum.

Die Ecdysozoa behoort tot Protostomia en is genoem na die algemene eienskap van groei deur vervelling.[88] Dit sluit die grootste dierefilum, die Arthropoda, in en bevat insekte, spinnekoppe, krappe, ens. Al dié diere se liggaam is in herhalende segmente verdeel wat gewoonlik in pare voorkom. Twee kleiner filums, Onychophora en Tardigrada, is nabye verwante en deel dié eienskappe. Ecdysozoa sluit ook die Nematoda, of rondewurms, in wat dalk die tweede grootste dierefilum is. Rondewurms is meestal mikroskopies en kom voor in feitlik elke omgewing waar daar water is.[89] ’n Paar is belangrike parasiete.[90] Kleiner filums wat aan hulle verwant is, is Nematomorpha, of die perdehaarwurms, Kinorhyncha, Priapulida en Loricifera. Hierdie groepe het ’n klein buikholte, wat ’n skynbuikholte genoem word.

 src=
Die wingerdslak Helix pomatia.

Lophotrochozoa

Lophotrochozoa, wat binne die groep Protostomia ontwikkel het, sluit twee van die mees suksesvolle dierefilums in, Mollusca en Annelida.[91][92] Eersgenoemde, wat die tweede grootste dierefilum is volgens spesies wat beskryf is, bevat diere soos slakke, mossels en pylinkvisse; laasgenoemde die gesegmenteerde wurms, soos erdwurms en bloedsuiers. Hierdie twee groepe is lank as nou verwant beskou omdat hulle soortgelyke larwes het, maar Annelida is as nader aan Arthropoda beskou omdat albei gesegmenteer is.[93] Nou word hulle beskou as die resultaat van saamlopende (maar onafhanklike) evolusie vanweë baie morfologiese en genetiese verskille tussen die twee filums.[94]

Lophotrochozoa sluit ook Nemertea, of die snoerwurms, Sipuncula en verskeie filums in wat ’n sirkel tentakels met trilhare om die mond het.[95]

Platyzoa sluit die filum Platyhelminth, of die platwurms, in.[96] Hulle is eers as van die mees primitiewe Bilateria beskou, maar dit lyk nou of hulle uit meer komplekse voorouers ontwikkel het.[97] Die groep bevat ’n paar parasiete, soos suigwurms and lintwurms.[96] Platwurms het nie ’n seloom of buikholte nie, nes hul naaste verwante, die mikroskopiese Gastrotricha.[98] Die ander Platyzoa-filums is meestal mikroskopies en het skynbuikholtes. Die belangrikste groep is Rotifera, of die raderdiertjies, wat algemeen in wateromgewings voorkom. Hulle sluit ook Acanthocephala, Gnathostomulida, Micrognathozoa en moontlik Cycliophora in.[99] Hierdie groepe het almal komplekse kake en behoort tot die filum Gnathifera.

Getal bestaande spesies

Diere kan in die algemeen in twee breë groepe verdeel word: gewerweldes (diere met ’n ruggraat) en ongewerweldes (sonder ’n ruggraat). Die helfte van alle gewerweldes wat beskryf is, is visse. Driekwart van alle ongewerweldes wat beskryf is, is insekte. Die volgende lys is van die bestaande spesies wat beskryf is vir elke groot subgroep van diere, soos beraam vir die IUBN-rooilys van bedreigde spesies, 2014.3.[100]

 src=
Die getal spesies volgens elke filum van die diereryk.

Meer as 95% van alle dierespesies in die wêreld wat beskryf is, is ongewerweldes.

Kladogram

Die Metazoa (diere) is verwant aan die swamme (Fungi) en het uit dieselfde groep eensellige eukariote ontstaan: die Ophistokonta. 'n Vereenvoudigde kladogram sien so uit:[101]

Opisthokonta

Fungi Fly agaric.svg


Holozoa

Ichthyospora


Filozoa?

Filasterea


Apoikozoa

Choanoflagellatea Cronoflagelado2.svg


Metazoa

Ctenophora Mertensia ovum.png



Porifera FMIB 49348 Ventriculites simplex, Toulmin Smith.jpeg




Placozoa Trichoplax mic-trans.png




Cnidaria 201208 Hydra.png



Bilateria Human.svg









Sien ook

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Eksterne skakels

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Dier: Brief Summary ( Afrikaans )

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Diere behoort tot Animalia, een van vyf (of soms ses) biologiese koninkryke. Dit word ook Metazoa genoem. Diere is oor die algemeen veelsellige, eukariotiese organismes wat ander organismes eet vir voeding. Die meeste diere kan in die een of ander stadium van hul lewe onafhanklik en spontaan beweeg.

Diere bestaan reeds van die Prekambrium (PreЄ) af, maar die meeste bekende dierefilums het tydens die Kambriese ontploffing, sowat 542 miljoen jaar gelede, in die fossielrekord verskyn. Diere kan oor die algemeen in gewerweldes en ongewerweldes verdeel word. Gewerweldes het ’n ruggraat en maak minder as 5% van alle dierespesies uit. Dit sluit in visse, amfibieë, reptiele, voëls en soogdiere. Die res is ongewerweldes; hulle het nie ’n ruggraat nie. Dit sluit in weekdiere (mossels, oesters, seekatte, pylinkvisse, slakke); geleedpotiges (insekte, spinnekoppe, skerpioene, krappe, krewe, garnale); ringwurms (erdwurms, bloedsuiers), rondewurms, platwurms, neteldiere (jellievisse, seeanemone, korale), ribkwalle en sponsdiere.

Die studie van diere word dierkunde of soölogie genoem.

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Animalia ( Asturian )

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Los animales (Animalia) o metazoos (Metazoa) constitúi un ampliu grupu d'organismos que son eucariotes, heterótrofos, pluricelulares y tisulares (sacante los poríferos). Caracterizase pola so amplia capacidá de movimientu, por non tener cloroplastu, nin paré celular, y pol so desenvolvimientu embrionariu; que traviesa una fase de blástula y determina un plan corporal fixu (anque munches especies puen sufrir una metamorfosis posterior). Animalia ye unu de los cinco reinos del dominiu Eukaryota, y a él pertenez el ser humanu.

Funciones

Alimentación

La mayoría de los animales nun pueden absorber comida; inxerir. Los animales evolucionaron de diverses formes p'alimentase. Los herbívoros comen plantes, los carnívoros comen otros animales; y los omnívoros aliméntense tantu de plantes como d'animales. Los detritívoros comen material vexetal y animal en descomposición. Los comedores por filtración son animales acuáticos que colen minúsculos organismos que llexen na agua. Los animales tamién formen rellaciones simbiótiques, nes que dos especies viven n'estrecha asociación mutua. Por casu un parásitu ye un tipu de simbionte que vive dientro o sobre otru organismu, el güéspede. El parásitu alimentar del güéspede y estropiar.

Respiración

No importa si viven na agua o na tierra, tolos animales alienden; esto significa que pueden tomar osíxenu y despidir dióxidu de carbonu. Gracies a los sos cuerpos bien simples y de delgaes parés, dellos animales utilicen l'espardimientu d'estes sustances al traviés de la piel. Sicasí, la mayoría de los animales evolucionaron complexos texíos y sistemes orgánicos pa la respiración.

​Circulación

Muchos animales acuáticos pequeños, como dellos viermes, utilicen solo l'espardimientu pa tresportar osíxenu y molécules de nutrientes a toles sos célules, y recoyer d'elles los productos de refugaya. L'espardimientu bastu porque estos animales apenes tienen una espesura d'unes cuantes célules. Sicasí, los animales más grandes tienen dalgún tipu de sistema circulatoriu pa mover sustances pel interior de los sos cuerpos.3​ExcreciónUn productu de refugaya primaria de les célules ye l'amoniacu, sustanza venenoso que contien nitróxenu. L'acumuladura d'amoniacu y otros productos de refugaya podríen matar a un animal. La mayoría de los animales tienen un sistema excretor que bien esanicia amoniacu o bien lo tresforma nuna sustanza menos tóxico que s'esanicia del cuerpu. Gracies a qu'esanicien les refugayes metabóliques, los sistemes excretores ayuden a caltener la homeóstasis. Los sistemes excretores varien, dende célules que bombien agua fora del cuerpu hasta órganos complexos como riñonos.

Respuesta

Los animales usen célules especializaes, llamaes célules nervioses, pa responder a los sucesos del so mediu ambiente. Na mayoría de los animales, les célules nervioses tán conectaes ente sigo pa formar un sistema nerviosu. Delles célules llamaes receptores, respuenden a soníos, lluz y otros estímulos esternos. Otres célules nervioses procesen información y determinen la respuesta del animal. La organización de les célules nervioses dientro del cuerpu camuda dramáticamente d'un fílum a otru.

Movimientu

Algunos animales adultos permanecen fixos nun sitiu. Anque munchos tienen movilidá. Sicasí tantu los fixos como los más rápidos de normal tienen músculos o texíos musculares que s'encurtien pa xenerar fuerza. La contracción muscular dexa que los animales movibles muévanse, de cutiu en combinación con una estructura llamada cadarma. Los músculos tamién ayuden a los animales, entá los más sedentarios, a comer y bombiar agua y otros líquidos fuera del cuerpu.

​Reproducción

La mayoría de los animales reprodúcense sexualmente por aciu la producción de gametos haploides. La reproducción sexual ayuda a crear y caltener la diversidá xenética d'una población. Poro, ayuda a ameyorar la capacidá d'una especie pa evolucionar colos cambeos del mediu ambiente. Munchos invertebraos tamién pueden reproducise asexualmente. La reproducción asexual da orixe a descendiente genéticamente idénticos a'l proxenitores. Esta forma de reproducción dexa que los animales aumenten rápido en cantidá.

Enllaces esternos

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Animalia: Brief Summary ( Asturian )

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... "Wikipedia:Wikipedia:Llista

Los animales (Animalia) o metazoos (Metazoa) constitúi un ampliu grupu d'organismos que son eucariotes, heterótrofos, pluricelulares y tisulares (sacante los poríferos). Caracterizase pola so amplia capacidá de movimientu, por non tener cloroplastu, nin paré celular, y pol so desenvolvimientu embrionariu; que traviesa una fase de blástula y determina un plan corporal fixu (anque munches especies puen sufrir una metamorfosis posterior). Animalia ye unu de los cinco reinos del dominiu Eukaryota, y a él pertenez el ser humanu.

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